LETTER 228. TO A.R. WALLACE. March 27th [1869].

I have lately (i.e., in new edition of the "Origin") (228/1. Fifth edition, 1869, pages 150-57.) been moderating my zeal, and attributing much more to mere useless variability. I did think I would send you the sheet, but I daresay you would not care to see it, in which I discuss Nageli's Essay on Natural Selection not affecting characters of no functional importance, and which yet are of high classificatory importance. Hooker is pretty well satisfied with what I have said on this head.

LETTER 229. TO J.D. HOOKER. Caerdeon, Barmouth, North Wales, July 24th [1869].

We shall be at home this day week, taking two days on the journey, and right glad I shall be. The whole has been a failure to me, but much enjoyment to the young...My wife has ailed a good deal nearly all the time; so that I loathe the place, with all its beauty. I was glad to hear what you thought of F. Muller, and I agree wholly with you. Your letter came at the nick of time, for I was writing on the very day to Muller, and I passed on your approbation of Chaps. X. and XI. Some time I should like to borrow the "Transactions of the New Zealand Institute," so as to read Colenso's article. (229/1. Colenso, "On the Maori Races of New Zealand." "N.Z. Inst. Trans." 1868, Pt. 3.) You must read Huxley v. Comte (229/2. "The Scientific Aspects of Positivism." "Fortnightly Review," 1869, page 652, and "Lay Sermons," 1870, page 162. This was a reply to Mr. Congreve's article, "Mr. Huxley on M. Comte," published in the April number of the "Fortnightly," page 407, which had been written in criticism of Huxley's article in the February number of the "Fortnightly," page 128, "On the Physical Basis of Life."); he never wrote anything so clever before, and has smashed everybody right and left in grand style. I had a vague wish to read Comte, and so had George, but he has entirely cured us of any such vain wish.

There is another article (229/3. "North British Review," Volume 50, 1869: "Geological Time," page 406. The papers reviewed are Sir William Thomson, "Trans. R. Soc. Edin." 1862; "Phil. Mag." 1863; Thomson and Tait, "Natural Philosophy," Volume I., App. D; Sir W. Thomson, "Proc. R. Soc. Edin." 1865; "Trans. Geol. Soc. Glasgow," 1868 and 1869; "Macmillan's Mag." 1862; Prof. Huxley, Presidential Address, "Geol. Soc. London," February, 1869; Dr. Hooker, Presidential Address, "Brit. Assoc." Norwich, 1868. Also the review on the "Origin" in the "North British Review," 1867, by Fleeming Jenkin, and an article in the "Pall Mall Gazette," May 3rd, 1869. The author treats the last-named with contempt as the work of an anonymous journalist, apparently unconscious of his own similar position.) just come out in last "North British," by some great mathematician, which is admirably done; he has a severe fling at you (229/4. The author of the "North British" article appears to us, at page 408, to misunderstand or misinterpret Sir J.D. Hooker's parable on "underpinning." See "Life and Letters," III., page 101 (note). Sir Joseph is attacked with quite unnecessary vehemence on another point at page 413.), but the article is directed against Huxley and for Thomson. This review shows me—not that I required being shown—how devilish a clever fellow Huxley is, for the reviewer cannot help admiring his abilities. There are some good specimens of mathematical arrogance in the review, and incidentally he shows how often astronomers have arrived at conclusions which are now seen to be mistaken; so that geologists might truly answer that we must be slow in admitting your conclusions. Nevertheless, all uniformitarians had better at once cry "peccavi,"—not but what I feel a conviction that the world will be found rather older than Thomson makes it, and far older than the reviewer makes it. I am glad I have faced and admitted the difficulty in the last edition of the "Origin," of which I suppose you received, according to order, a copy.

LETTER 230. TO J.D. HOOKER. Down, August 7th [1869].

There never was such a good man as you for telling me things which I like to hear. I am not at all surprised that Hallett has found some varieties of wheat could not be improved in certain desirable qualities as quickly as at first. All experience shows this with animals; but it would, I think, be rash to assume, judging from actual experience, that a little more improvement could not be got in the course of a century, and theoretically very improbable that after a few thousands [of years] rest there would not be a start in the same line of variation. What astonishes me as against experience, and what I cannot believe, is that varieties already improved or modified do not vary in other respects. I think he must have generalised from two or three spontaneously fixed varieties. Even in seedlings from the same capsule some vary much more than others; so it is with sub-varieties and varieties. (230/1. In a letter of August 13th, 1869, Sir J.D. Hooker wrote correcting Mr. Darwin's impression: "I did not mean to imply that Hallett affirmed that all variation stopped—far from it: he maintained the contrary, but if I understand him aright, he soon arrives at a point beyond which any further accumulation in the direction sought is so small and so slow that practically a fixity of type (not absolute fixity, however) is the result.")

It is a grand fact about Anoplotherium (230/2. This perhaps refers to the existence of Anoplotherium in the S. American Eocene formation: it is one of the points in which the fauna of S. America resembles Europe rather than N. America. (See Wallace "Geographical Distribution," I., page 148.)), and shows how even terrestrial quadrupeds had time formerly to spread to very distinct regions. At each epoch the world tends to get peopled pretty uniformly, which is a blessing for Geology.

The article in "N. British Review" (230/3. See Letter 229.) is well worth reading scientifically; George D. and Erasmus were delighted with it. How the author does hit! It was a euphuism to speak of a fling at you: it was a kick. He is very unfair to Huxley, and accuses him of "quibbling," etc.; yet the author cannot help admiring him extremely. I know I felt very small when I finished the article. You will be amused to observe that geologists have all been misled by Playfair, who was misled by two of the greatest mathematicians! And there are other such cases; so we could turn round and show your reviewer how cautious geologists ought to be in trusting mathematicians.

There is another excellent original article, I feel sure by McClennan, on Primeval Man, well worth reading.

I do not quite agree about Sabine: he is unlike every other soldier or sailor I ever heard of if he would not put his second leg into the tomb with more satisfaction as K.C.B. than as a simple man. I quite agree that the Government ought to have made him long ago, but what does the Government know or care for Science? So much for your splenditious letter.

LETTER 231. TO J.D. HOOKER. Down, August 14th [1869?]

I write one line to tell you that you are a real good man to propose coming here for a Sunday after Exeter. Do keep to this good intention...I am sure Exeter and your other visit will do you good. I often wonder how you stand all your multifarious work.

I quite agree about the folly of the endless subscriptions for dead men; but Faraday is an exception, and if you will pay three guineas for me, it will save me some trouble; but it will be best to enclose a cheque, which, as you will see, must be endorsed. If you read the "North British Review," you will like to know that George has convinced me, from correspondence in style, and spirit, that the article is by Tait, the co-worker with Thomson.

I was much surprised at the leaves of Drosophyllum being always rolled backwards at their tips, but did not know that it was a unique character.

(PLATE: SIR J.D. HOOKER, 1870? From a photograph by Wallich.)

LETTER 232. TO J.D. HOOKER. Down, November 13th [1869].

I heard yesterday from a relation who had seen in a newspaper that you were C.B. I must write one line to say "Hurrah," though I wish it had been K.C.B., as it assuredly ought to have been; but I suppose they look at K.C.B. before C.B. as a dukedom before an earldom.

We had a very successful week in London, and I was unusually well and saw a good many persons, which, when well, is a great pleasure to me. I had a jolly talk with Huxley, amongst others. And now I am at the same work as before, and shall be for another two months—namely, putting ugly sentences rather straighter; and I am sick of the work, and, as the subject is all on sexual selection, I am weary of everlasting males and females, cocks and hens.

It is a shame to bother you, but I should like some time to hear about the C.B. affair.

I have read one or two interesting brochures lately—viz., Stirling the Hegelian versus Huxley and protoplasm; Tylor in "Journal of Royal Institute" on the survivals of old thought in modern civilisation.

Farewell. I am as dull as a duck, both male and female.

To Dr. Hooker, C.B., F.R.S.

Dr. Hooker, K.C.B. (This looks better).

P.S. I hear a good account of Bentham's last address (232/1. Presidential Address, chiefly on Geographical Distribution, delivered before the "Linn. Soc." May 24th, 1869.), which I am now going to read.

I find that I have blundered about Bentham's address. Lyell was speaking about one that I read some months ago; but I read half of it again last night, and shall finish it. Some passages are either new or were not studied enough by me before. It strikes me as admirable, as it did on the first reading, though I differ in some few points.

Such an address is worth its weight in gold, I should think, in making converts to our views. Lyell tells me that Bunbury has been wonderfully impressed with it, and he never before thought anything of our views on evolution.

P.S. (2). I have just read, and like very much, your review of Schimper. (232/2. A review of Schimper's "Traite de Paleontologie Vegetale," the first portion of which was published in 1869. "Nature," November 11th, 1869, page 48.)

LETTER 233. TO J.D. HOOKER. Down, November 19th [1869].

Thank you much for telling me all about the C.B., for I much wished to hear. It pleases me extremely that the Government have done this much; and as the K.C.B.'s are limited in number (which I did not know), I excuse it. I will not mention what you have told me to any one, as it would be Murchisonian. But what a shame it is to use this expression, for I fully believe that Murchison would take any trouble to get any token of honour for any man of science.

I like all scientific periodicals, including poor "Scientific Opinion," and I think higher than you do of "Nature." Lord, what a rhapsody that was of Goethe, but how well translated; it seemed to me, as I told Huxley, as if written by the maddest English scholar. It is poetry, and can I say anything more severe? The last number of the "Academy" was splendid, and I hope it will soon come out fortnightly. I wish "Nature" would search more carefully all foreign journals and transactions.

I am now reading a German thick pamphlet (233/1. "Die Abhangigheit der Pflanzengestalt von Klima und Boden. Ein Beitrag zur Lehre von der Enstehung und Verbreitung der Arten, etc." Festschrift zur 43 Versammlung Deutscher Naturforscher und Aertze in Innsbruck (Innsbruck, 1869).) by Kerner on Tubocytisus; if you come across it, look at the map of the distribution of the eighteen quasi-species, and at the genealogical tree. If the latter, as the author says, was constructed solely from the affinities of the forms, then the distribution is wonderfully interesting; we may see the very steps of the formation of a species. If you study the genealogical tree and map, you will almost understand the book. The two old parent connecting links just keep alive in two or three areas; then we have four widely extended species, their descendants; and from them little groups of newer descendants inhabiting rather small areas...

LETTER 234. TO CAMILLE DARESTE. Down, November 20th, 1869.

Dear Sir,

I am glad that you are a candidate for the Chair of Physiology in Paris. As you are aware from my published works, I have always considered your investigations on the production of monstrosities as full of interest. No subject is at the present time more important, as far as my judgment goes, than the ascertaining by experiment how far structure can be modified by the direct action of changed conditions; and you have thrown much light on this subject.

I observe that several naturalists in various parts of Europe have lately maintained that it is now of the highest interest for science to endeavour to lessen, as far as possible, our profound ignorance on the cause of each individual variation; and, as Is. Geoffroy St. Hilaire long ago remarked, monstrosities cannot be separated by any distinct line from slighter variations.

With my best wishes for your success in obtaining the Professorship, and with sincere respect.

I have the honour to remain, dear sir, Yours faithfully, CHARLES DARWIN.



CHAPTER 1.V.—EVOLUTION, 1870-1882.

LETTER 235. TO J. JENNER WEIR. Down, March 17th [1870].

It is my decided opinion that you ought to send an account to some scientific society, and I think to the Royal Society. (235/1. Mr. Jenner Weir's case is given in "Animals and Plants," Edition II., Volume I., page 435, and does not appear to have been published elsewhere. The facts are briefly that a horse, the offspring of a mare of Lord Mostyn's, which had previously borne a foal by a quagga, showed a number of quagga-like characters, such as stripes, low-growing mane, and elongated hoofs. The passage in "Animals and Plants," to which he directs Mr. Weir's attention in reference to Carpenter's objection, is in Edition I., Volume I., page 405: "It is a most improbable hypothesis that the mere blood of one individual should affect the reproductive organs of another individual in such a manner as to modify the subsequent offspring. The analogy from the direct action of foreign pollen on the ovarium and seed-coats of the mother plant strongly supports the belief that the male element acts directly on the reproductive organs of the female, wonderful as is this action, and not through the intervention of the crossed embryo." For references to Mr. Galton's experiments on transfusion of blood, see Letter 273.) I would communicate it if you so decide. You might give as a preliminary reason the publication in the "Transactions" of the celebrated Morton case and the pig case by Mr. Giles. You might also allude to the evident physiological importance of such facts as bearing on the theory of generation. Whether it would be prudent to allude to despised pangenesis I cannot say, but I fully believe pangenesis will have its successful day. Pray ascertain carefully the colour of the dam and sire. See about duns in my book ["Animals and Plants"], Volume I., page 55. The extension of the mane and form of hoofs are grand new facts. Is the hair of your horse at all curly? for [an] observed case [is] given by me (Volume II., page 325) from Azara of correlation of forms of hoof with curly hairs. See also in my book (Volume I., page 55; Volume II., page 41) how exceedingly rare stripes are on the faces of horses in England. Give the age of your horse.

You are aware that Dr. Carpenter and others have tried to account for the effects of a first impregnation from the influence of the blood of the crossed embryo; but with physiologists who believe that the reproductive elements are actually formed by the reproductive glands, this view is inconsistent. Pray look at what I have said in "Domestic Animals" (Volume I., pages 402-5) against this doctrine. It seems to me more probable that the gemmules affect the ovaria alone. I remember formerly speculating, like you, on the assertion that wives grow like their husbands; but how impossible to eliminate effects of imitation and same habits of life, etc. Your letter has interested me profoundly.

P.S.—Since publishing I have heard of additional cases—a very good one in regard to Westphalian pigs crossed by English boar, and all subsequent offspring affected, given in "Illust. Landwirth-Zeitung," 1868, page 143.

I have shown that mules are often striped, though neither parent may be striped,—due to ancient reversion. Now, Fritz Muller writes to me from S. Brazil: "I have been assured, by persons who certainly never had heard of Lord Morton's mare, that mares which have borne hybrids to an ass are particularly liable to produce afterwards striped ass-colts." So a previous fertilisation apparently gives to the subsequent offspring a tendency to certain characters, as well as characters actually possessed by the first male.

In the reprint (not called a second edition) of my "Domestic Animals" I give a good additional case of subsequent progeny of hairless dog being hairy from effects of first impregnation.

P.S. 2nd. The suggestion, no doubt, is superfluous, but you ought, I think, to measure extension of mane beyond a line joining front or back of ears, and compare with horse. Also the measure (and give comparison with horse), length, breadth, and depth of hoofs.

LETTER 236. TO J.D. HOOKER. Down, July 12th [1870].

Your conclusion that all speculation about preordination is idle waste of time is the only wise one; but how difficult it is not to speculate! My theology is a simple muddle; I cannot look at the universe as the result of blind chance, yet I can see no evidence of beneficent design or indeed of design of any kind, in the details. As for each variation that has ever occurred having been preordained for a special end, I can no more believe in it than that the spot on which each drop of rain falls has been specially ordained.

Spontaneous generation seems almost as great a puzzle as preordination. I cannot persuade myself that such a multiplicity of organisms can have been produced, like crystals, in Bastian's (236/1. On September 2nd, 1872, Mr. Darwin wrote to Mr. Wallace, in reference to the latter's review of "The Beginnings of Life," by H.C. Bastian (1872), in "Nature," 1872, pages 284-99: "At present I should prefer any mad hypothesis, such as that every disintegrated molecule of the lowest forms can reproduce the parent-form; and that these molecules are universally distributed, and that they do not lose their vital power until heated to such a temperature that they decompose like dead organic particles.") solutions of the same kind. I am astonished that, as yet, I have met with no allusion to Wyman's positive statement (236/2. "Observations and Experiments on Living Organisms in Heated Water," by Jeffries Wyman, Prof. of Anatomy, Harvard Coll. ("Amer. Journ. Sci." XLIV., 1867, page 152.) Solutions of organic matter in hermetically sealed flasks were immersed in boiling water for various periods. "No infusoria of any kind appeared if the boiling was prolonged beyond a period of five hours.") that if the solutions are boiled for five hours no organisms appear; yet, if my memory serves me, the solutions when opened to air immediately became stocked. Against all evidence, I cannot avoid suspecting that organic particles (my "gemmules" from the separate cells of the lower creatures!) will keep alive and afterwards multiply under proper conditions.

What an interesting problem it is.

LETTER 237. TO W.B. TEGETMEIER. Down, July 15th [1870].

It is very long since I have heard from you, and I am much obliged for your letter. It is good news that you are going to bring out a new edition of your Poultry book (237/1. "The Poultry Book," 1872.), and you are quite at liberty to use all my materials. Thanks for the curious case of the wild duck variation: I have heard of other instances of a tendency to vary in one out of a large litter or family. I have too many things in hand at present to profit by your offer of the loan of the American Poultry book.

Pray keep firm to your idea of working out the subject of analogous variations (237/2. "By this term I mean that similar characters occasionally make their appearance in the several varieties or races descended from the same species, and more rarely in the offspring of widely distinct species" ("Animals and Plants," II., Edition II., page 340).) with pigeons; I really think you might thus make a novel and valuable contribution to science. I can, however, quite understand how much your time must be occupied with the never-ending, always-beginning editorial cares.

I keep much as usual, and crawl on with my work.

LETTER 238. TO J.D. HOOKER. Down, September 27th [1870].

Yours was a splendid letter, and I was very curious to hear something about the Liverpool meeting (238/1. Mr. Huxley was President of the British Association at Liverpool in 1870. His Presidential Address on "Biogenesis and Abiogenesis" is reprinted in his collected Essays, VIII., page 229. Some account of the meeting is given in Huxley's "Life and Letters," Volume I., pages 332, 336.), which I much wished to be successful for Huxley's sake. I am surprised that you think his address would not have been clear to the public; it seemed to me as clear as water. The general line of his argument might have been answered by the case of spontaneous combustion: tens of thousands of cases of things having been seen to be set on fire would be no true argument against any one who maintained that flames sometimes spontaneously burst forth. I am delighted at the apotheosis of Sir Roderick; I can fancy what neat and appropriate speeches he would make to each nobleman as he entered the gates of heaven. You ask what I think about Tyndall's lecture (238/2. Tyndall's lecture was "On the Scientific Uses of the Imagination."): it seemed to me grand and very interesting, though I could not from ignorance quite follow some parts, and I longed to tell him how immensely it would have been improved if all the first part had been made very much less egotistical. George independently arrived at the same conclusion, and liked all the latter part extremely. He thought the first part not only egotistical, but rather clap-trap.

How well Tyndall puts the "as if" manner of philosophising, and shows that it is justifiable. Some of those confounded Frenchmen have lately been pitching into me for using this form of proof or argument.

I have just read Rolleston's address in "Nature" (238/3. Presidential Address to the Biological Section, British Association, 1870. "Nature," September 22nd, 1870, page 423. Rolleston referred to the vitality of seeds in soil, a subject on which Darwin made occasional observations. See "Life and Letters," II., page 65.): his style is quite unparalleled! I see he quotes you about seed, so yesterday I went and observed more carefully the case given in the enclosed paper, which perhaps you might like to read and burn.

How true and good what you say about Lyell. He is always the same; Dohrn was here yesterday, and was remarking that no one stood higher in the public estimation of Germany than Lyell.

I am truly and profoundly glad that you are thinking of some general work on Geographical Distribution, or so forth; I hope to God that your incessant occupations may not interrupt this intention. As for my book, I shall not have done the accursed proofs till the end of November (238/4. The proofs of the "Descent of Man" were finished on January 15th, 1871.): good Lord, what a muddled head I have got on my wretched old shoulders.

LETTER 239. TO H. SETTEGAST. Down, September 29th, 1870.

I am very much obliged for your kind letter and present of your beautiful volume. (239/1. "Die Thierzucht," 1868.) Your work is not new to me, for I heard it so highly spoken of that I procured a copy of the first edition. It was a great gratification to me to find a man who had long studied with a philosophical spirit our domesticated animals, and who was highly competent to judge, agreeing to a large extent with my views. I regretted much that I had not known your work when I published my last volumes.

I am surprised and pleased to hear that science is not quite forgotten under the present exciting state of affairs. Every one whom I know in England is an enthusiastic wisher for the full and complete success of Germany.

P.S. I will give one of my two copies of your work to some public scientific library in London.

LETTER 240. TO THE EDITOR OF THE "PALL MALL GAZETTE." Down, March 24th [1871].

Mr. Darwin presents his compliments to the Editor, and would be greatly obliged if he would address and post the enclosed letter to the author of the two admirable reviews of the "Descent of Man." (240/1. The notices of the "Descent of Man," published in the "Pall Mall Gazette" of March 20th and 21st, 1871, were by Mr. John Morley. We are indebted to the Editor of the "Pall Mall Gazette" for kindly allowing us to consult his file of the journal.)

LETTER 241. TO JOHN MORLEY. Down, March 24th, 1871.

From the spirit of your review in the "Pall Mall Gazette" of my last book, which has given me great pleasure, I have thought that you would perhaps inform me on one point, withholding, if you please, your name.

You say that my phraseology on beauty is "loose scientifically, and philosophically most misleading." (241/1. "Mr. Darwin's work is one of those rare and capital achievements of intellect which effect a grave modification throughout all the highest departments of the realm of opinion...There is throughout the description and examination of Sexual Selection a way of speaking of beauty, which seems to us to be highly unphilosophical, because it assumes a certain theory of beauty, which the most competent modern thinkers are too far from accepting, to allow its assumption to be quite judicious...Why should we only find the aesthetic quality in birds wonderful, when it happens to coincide with our own? In other words, why attribute to them conscious aesthetic qualities at all? There is no more positive reason for attributing aesthetic consciousness to the Argus pheasant than there is for attributing to bees geometric consciousness of the hexagonal prisms and rhombic plates of the hive which they so marvellously construct. Hence the phraseology which Mr. Darwin employs in this part of the subject, though not affecting the degree of probability which may belong to this theory, seems to us to be very loose scientifically, and philosophically most misleading."—"Pall Mall Gazette.") This is not at all improbable, as it is almost a lifetime since I attended to the philosophy of aesthetics, and did not then think that I should ever make use of my conclusions. Can you refer me to any one or two books (for my power of reading is not great) which would illumine me? or can you explain in one or two sentences how I err? Perhaps it would be best for me to explain what I mean by the sense of beauty in its lowest stage of development, and which can only apply to animals. When an intense colour, or two tints in harmony, or a recurrent and symmetrical figure please the eye, or a single sweet note pleases the ear, I call this a sense of beauty; and with this meaning I have spoken (though I now see in not a sufficiently guarded manner) of a taste for the beautiful being the same in mankind (for all savages admire bits of bright cloth, beads, plumes, etc.) and in the lower animals. If the blue and yellow plumage of a macaw (241/2. "What man deems the horrible contrasts of yellow and blue attract the macaw, while ball-and-socket-plumage attracts the Argus pheasant"—"Pall Mall Gazette," March 21st, 1871, page 1075.) pleases the eye of this bird, I should say that it had a sense of beauty, although its taste was bad according to our standard. Now, will you have the kindness to tell me how I can learn to see the error of my ways? Of course I recognise, as indeed I have remarked in my book, that the sense of beauty in the case of scenery, pictures, etc., is something infinitely complex, depending on varied associations and culture of the mind. From a very interesting review in the "Spectator," and from your and Wallace's review, I perceive that I have made a great oversight in not having said what little I could on the acquisition of the sense for the beautiful by man and the lower animals. It would indeed be an immense advantage to an author if he could read such criticisms as yours before publishing. At page 11 of your review you accidentally misquote my words placed by you within inverted commas, from my Volume II., page 354: I say that "man cannot endure any great change," and the omitted words "any great" make all the difference in the discussion. (241/3. "Mr. Darwin tells us, and gives us excellent reasons for thinking, that 'the men of each race prefer what they are accustomed to behold; they cannot endure change.' Yet is there not an inconsistency between this fact and the other that one race differs from another exactly because novelties presented themselves, and were eagerly seized and propagated?")

Permit me to add a few other remarks. I believe your criticism is quite just about my deficient historic spirit, for I am aware of my ignorance in this line. (241/4. "In the historic spirit, however, Mr. Darwin must fairly be pronounced deficient. When, for instance, he speaks of the 'great sin of slavery' having been general among primitive nations, he forgets that, though to hold a slave would be a sinful degradation to a European to-day, the practice of turning prisoners of war into slaves, instead of butchering them, was not a sin at all, but marked a decided improvement in human manners.") On the other hand, if you should ever be led to read again Chapter III., and especially Chapter V., I think you will find that I am not amenable to all your strictures; though I felt that I was walking on a path unknown to me and full of pitfalls; but I had the advantage of previous discussions by able men. I tried to say most emphatically that a great philosopher, law-giver, etc., did far more for the progress of mankind by his writings or his example than by leaving a numerous offspring. I have endeavoured to show how the struggle for existence between tribe and tribe depends on an advance in the moral and intellectual qualities of the members, and not merely on their capacity of obtaining food. When I speak of the necessity of a struggle for existence in order that mankind should advance still higher in the scale, I do not refer to the MOST, but "to the MORE highly gifted men" being successful in the battle for life; I referred to my supposition of the men in any country being divided into two equal bodies—viz., the more and the less highly gifted, and to the former on an average succeeding best.

But I have much cause to apologise for the length of this ill-expressed letter. My sole excuse is the extraordinary interest which I have felt in your review, and the pleasure which I have experienced in observing the points which have attracted your attention. I must say one word more. Having kept the subject of sexual selection in my mind for very many years, and having become more and more satisfied with it, I feel great confidence that as soon as the notion is rendered familiar to others, it will be accepted, at least to a much greater extent than at present. With sincere respect and thanks...

LETTER 242. TO JOHN MORLEY. Down, April 14th [1871].

As this note requires no answer, I do not scruple to write a few lines to say how faithful and full a resume you have given of my notions on the moral sense in the "Pall Mall," and to make a few extenuating or explanatory remarks. (242/1. "What is called the question of the moral sense is really two: how the moral faculty is acquired, and how it is regulated. Why do we obey conscience or feel pain in disobeying it? And why does conscience prescribe one kind of action and condemn another kind? To put it more technically, there is the question of the subjective existence of conscience, and there is the question of its objective prescriptions. First, why do I think it obligatory to do my duty? Second, why do I think it my duty to do this and not do that? Although, however, the second question ought to be treated independently, for reasons which we shall presently suggest, the historical answer to it, or the various grounds on which men have identified certain sorts of conduct with duty, rather than conduct of the opposite sorts, throws light on the other question of the conditions of growth of the idea of duty as a sovereign and imperial director. Mr. Darwin seems to us not to have perfectly recognised the logical separation between the two sides of the moral sense question. For example, he says (i. 97) that 'philosophers of the derivative school of morals formerly assumed that the foundation of morality lay in a form of Selfishness; but more recently in the Greatest Happiness principle.' But Mr. Mill, to whom Mr. Darwin refers, has expressly shown that the Greatest Happiness principle is a STANDARD, and not a FOUNDATION, and that its validity as a standard of right and wrong action is just as tenable by one who believes the moral sense to be innate, as by one who holds that it is acquired. He says distinctly that the social feelings of mankind form 'the natural basis of sentiment for utilitarian morality.' So far from holding the Greatest Happiness principle to be the foundation of morality, he would describe it as the forming principle of the superstructure of which the social feelings of mankind are the foundation. Between Mr. Darwin and utilitarians, as utilitarians, there is no such quarrel as he would appear to suppose. The narrowest utilitarian could say little more than Mr. Darwin says (ii. 393): 'As all men desire their own happiness, praise or blame is bestowed on actions and motives according as they tend to this end; and, as happiness is an essential part of the general good, the Greatest Happiness principle INDIRECTLY serves as a NEARLY safe standard of right and wrong.' It is perhaps not impertinent to suspect that the faltering adverbs which we have printed in italics indicate no more than the reluctance of a half-conscious convert to pure utilitarianism. In another place (i. 98) he admits that 'as all wish for happiness, the Greatest Happiness principle will have become a most important secondary guide and object, the social instincts, including sympathy, always serving as the primary impulse and guide.' This is just what Mr. Mill says, only instead of calling the principle a secondary guide, he would call it a standard, to distinguish it from the social impulse, in which, as much as Mr. Darwin, he recognises the base and foundation."—"Pall Mall Gazette," April 12th, 1871.) How the mistake which I have made in speaking of greatest happiness as the foundation of morals arose, is utterly unintelligible to me: any time during the last several years I should have laughed such an idea to scorn. Mr. Lecky never made a greater blunder, and your kindness has made you let me off too easily. (242/2. In the first edition of the "Descent of Man," I., page 97, Mr. Lecky is quoted as one of those who assumed that the "foundation of morality lay in a form of selfishness; but more recently in the 'greatest happiness' principle." Mr. Lecky's name is omitted in this connection in the second edition, page 120. In this edition Mr. Darwin makes it clearer that he attaches most importance to the social instinct as the "primary impulse and guide.") With respect to Mr. Mill, nothing would have pleased me more than to have relied on his great authority with respect to the social instincts, but the sentence which I quote at [Volume I.] page 71 ("if, as is my own belief, the moral feelings are not innate, but acquired, they are not for that reason less natural") seems to me somewhat contradictory with the other words which I quote, so that I did not know what to think; more especially as he says so very little about the social instincts. When I speak of intellectual activity as the secondary basis of conscience, I meant in my own mind secondary in period of development; but no one could be expected to understand so great an ellipse. With reference to your last sentence, do you not think that man might have retrograded in his parental, marriage, and other instincts without having retrograded in his social instincts? and I do not think that there is any evidence that man ever existed as a non-social animal. I must add that I have been very glad to read your remarks on the supposed case of the hive-bee: it affords an amusing contrast with what Miss Cobbe has written in the "Theological Review." (242/3. Mr. Darwin says ("Descent of Man" Edition I., Volume I., page 73; Edition II., page 99), "that if men lived like bees our unmarried females would think it a sacred duty to kill their brothers." Miss Cobbe remarks on this "that the principles of social duty would be reversed" ("Theological Review," April 1872). Mr. Morley, on the other hand, says of Darwin's assertion, that it is "as reassuring as the most absolute of moralists could desire. For it is tantamount to saying that the foundations of morality, the distinctions of right and wrong, are deeply laid in the very conditions of social existence; that there is in face of these conditions a positive and definite difference between the moral and the immoral, the virtuous and the vicious, the right and the wrong, in the actions of individuals partaking of that social existence.") Undoubtedly the great principle of acting for the good of all the members of the same community, and therefore the good of the species, would still have held sovereign sway.

LETTER 243. TO J.D. HOOKER.

(243/1. Sir Joseph Hooker wrote (August 5th, 1871) to Darwin about Lord Kelvin's Presidential Address at the Edinburgh meeting of the British Association: "It seems to me to be very able indeed; and what a good notion it gives of the gigantic achievement of mathematicians and physicists!—it really made one giddy to read of them. I do not think Huxley will thank him for his reference to him as a positive unbeliever in spontaneous generation—these mathematicians do not seem to me to distinguish between un-belief and a-belief. I know no other name for the state of mind that is produced under the term scepticism. I had no idea before that pure Mathematics had achieved such wonders in practical science. The total absence of any allusion to Tyndall's labours, even when comets are his theme, seems strange to me.")

Haredene, Albury, Guildford, August 6th [1871].

I have read with greatest interest Thomson's address; but you say so EXACTLY AND FULLY all that I think, that you have taken all the words from my mouth; even about Tyndall. It is a gain that so wonderful a man, though no naturalist, should become a convert to evolution; Huxley, it seems, remarked in his speech to this effect. I should like to know what he means about design,—I cannot in the least understand, for I presume he does not believe in special interpositions. (243/2. See "British Association Report," page cv. Lord Kelvin speaks very doubtfully of evolution. After quoting the concluding passage of the "Origin," he goes on, "I have omitted two sentences...describing briefly the hypothesis of 'the origin of species by Natural Selection,' because I have always felt that this hypothesis does not contain the true theory of evolution, IF EVOLUTION THERE HAS BEEN in biology" (the italics are not in the original). Lord Kelvin then describes as a "most valuable and instructive criticism," Sir John Herschel's remark that the doctrine of Natural Selection is "too like the Laputan method of making books, and that it did not sufficiently take into account a continually guiding and controlling intelligence." But it should be remembered that it was in this address of Lord Kelvin's that he suggested the possibility of "seed-bearing meteoric stones moving about through space" inoculating the earth with living organisms; and if he assumes that the whole population of the globe is to be traced back to these "moss-grown fragments from the ruins of another world," it is obvious that he believes in a form of evolution, and one in which a controlling intelligence is not very obvious, at all events not in the initial and all-important stage.) Herschel's was a good sneer. It made me put in the simile about Raphael's Madonna, when describing in the "Descent of Man" the manner of formation of the wondrous ball-and-socket ornaments, and I will swear to the truth of this case. (243/3. See "Descent of Man," II., page 141. Darwin says that no one will attribute the shading of the "eyes" on the wings of the Argus pheasant to the "fortuitous concourse of atoms of colouring-matter." He goes on to say that the development of the ball-and-socket effect by means of Natural Selection seems at first as incredible as that "one of Raphael's Madonnas should have been formed by the selection of chance daubs of paint." The remark of Herschel's, quoted in "Life and Letters," II., page 241, that the "Origin" illustrates the "law of higgledy-piggledy," is probably a conversational variant of the Laputan comparison which gave rise to the passage in the "Descent of Man" (see Letter 130).)

You know the oak-leaved variety of the common honeysuckle; I could not persuade a lady that this was not the result of the honeysuckle climbing up a young oak tree! Is this not like the Viola case?

LETTER 244. TO JOHN LUBBOCK (LORD AVEBURY). Haredene, Albury, Guildford, August 12th [1871].

I hope the proof-sheets having been sent here will not inconvenience you. I have read them with infinite satisfaction, and the whole discussion strikes me as admirable. I have no books here, and wish much I could see a plate of Campodea. (244/1. "On the Origin of Insects." By Sir John Lubbock, Bart. "Journ. Linn. Soc. (Zoology)," Volume XI., 1873, pages 422-6. (Read November 2nd, 1871.) In the concluding paragraph the author writes, "If these views are correct the genus Campodea [a beetle] must be regarded as a form of remarkable interest, since it is the living representative of a primaeval type from which not only the Collembola and Thysanura, but the other great orders of insects, have all derived their origin." (See also "Brit. Assoc. Report," 1872, page 125—Address by Sir John Lubbock; and for a figure of Campodea see "Nature," Volume VII., 1873, page 447.) I never reflected much on the difficulty which you indicate, and on which you throw so much light. (244/2. The difficulty alluded to is explained by the first sentence of Lord Avebury's paper. "The Metamorphoses of this group (Insects) have always seemed to me one of the greatest difficulties of the Darwinian theory...I feel great difficulty in conceiving by what natural process an insect with a suctorial mouth, like that of a gnat or butterfly, could be developed from a powerfully mandibulate type like the orthoptera, or even from the neuroptera...A clue to the difficulty may, I think, be found in the distinction between the developmental and adaptive changes to which I called the attention of the Society in a previous memoir."

The distinction between developmental and adaptive changes is mentioned, but not discussed, in the paper "On the Origin of Insects" (loc. cit., page 422); in a former paper, "On the Development of Chloeon (Ephemera) dimidiatum ("Trans. Linn. Soc." XXV. page 477, 1866), this question is dealt with at length.) I have only a few trifling remarks to make. At page 44 I wish you had enlarged a little on what you have said of the distinction between developmental and adaptive changes; for I cannot quite remember the point, and others will perhaps be in the same predicament. I think I always saw that the larva and the adult might be separately modified to any extent. Bearing in mind what strange changes of function parts undergo, with the intermediate state of use (244/3. This slightly obscure phrase may be paraphrased, "the gradational stages being of service to the organism."), it seems to me that you speak rather too boldly on the impossibility of a mandibulate insect being converted into a sucking insect (244/4. "There are, however, peculiar difficulties in those cases in which, as among the lepidoptera, the same species is mandibulate as a larva and suctorial as an embryo" (Lubbock, "Origin of Insects," page 423).); not that I in the least doubt the value of your explanation.

Cirripedes passing through what I have called a pupal state (244/5. "Hence, the larva in this, its last stage, cannot eat; it may be called a "locomotive Pupa;" its whole organisation is apparently adapted for the one great end of finding a proper site for its attachment and final metamorphosis." ("A Monograph on the Sub-Class Cirripedia." By Charles Darwin. London, Ray Soc., 1851.)) so far as their mouths are concerned, rather supports what you say at page 52.

At page 40 your remarks on the Argus pheasant (244/6. There is no mention of the Argus pheasant in the published paper.) (though I have not the least objection to them) do not seem to me very appropriate as being related to the mental faculties. If you can spare me these proof-sheets when done with, I shall be obliged, as I shall be correcting a new edition of the "Origin" when I return home, though this subject is too large for me to enter on. I thank you sincerely for the great interest which your discussion has given me.

LETTER 245. TO J.D. HOOKER.

(245/1. The following letter refers to Mivart's "Genesis of Species.")

Down, September 16th [1871].

I am preparing a new and cheap edition of the "Origin," and shall introduce a new chapter on gradation, and on the uses of initial commencements of useful structures; for this, I observe, has produced the greatest effect on most persons. Every one of his [Mivart's] cases, as it seems to me, can be answered in a fairly satisfactory manner. He is very unfair, and never says what he must have known could be said on my side. He ignores the effect of use, and what I have said in all my later books and editions on the direct effects of the conditions of life and so-called spontaneous variation. I send you by this post a very clever, but ill-written review from N. America by a friend of Asa Gray, which I have republished. (245/2. Chauncey Wright in the "North American Review," Volume CXIII., reprinted by Darwin and published as a pamphlet (see "Life and Letters," III., page 145).)

I am glad to hear about Huxley. You never read such strong letters Mivart wrote to me about respect towards me, begging that I would call on him, etc., etc.; yet in the "Q. Review" (245/3. See "Quarterly Review," July 1871; also "Life and Letters," III., page 147.) he shows the greatest scorn and animosity towards me, and with uncommon cleverness says all that is most disagreeable. He makes me the most arrogant, odious beast that ever lived. I cannot understand him; I suppose that accursed religious bigotry is at the root of it. Of course he is quite at liberty to scorn and hate me, but why take such trouble to express something more than friendship? It has mortified me a good deal.

LETTER 246. TO J.D. HOOKER. Down, October 4th [1871].

I am quite delighted that you think so highly of Huxley's article. (246/1. A review of Wallace's "Natural Selection," of Mivart's "Genesis of Species," and of the "Quarterly Review" article on the "Descent of Man" (July, 1871), published in the "Contemporary Review" (1871), and in Huxley's "Collected Essays," II., page 120.) I was afraid of saying all I thought about it, as nothing is so likely as to make anything appear flat. I thought of, and quite agreed with, your former saying that Huxley makes one feel quite infantile in intellect. He always thus acts on me. I exactly agree with what you say on the several points in the article, and I piled climax on climax of admiration in my letter to him. I am not so good a Christian as you think me, for I did enjoy my revenge on Mivart. He (i.e. Mivart) has just written to me as cool as a cucumber, hoping my health is better, etc. My head, by the way, plagues me terribly, and I have it light and rocking half the day. Farewell, dear old friend—my best of friends.

LETTER 247. TO JOHN FISKE.

(247/1. Mr. Fiske, who is perhaps best known in England as the author of "Outlines of Cosmic Philosophy," had sent to Mr. Darwin some reports of the lectures given at Harvard University. The point referred to in the postscript in Mr. Darwin's letter is explained by the following extract from Mr. Fiske's work: "I have endeavoured to show that the transition from animality (or bestiality, stripping the word of its bad connotations) to humanity must have been mainly determined by the prolongation of infancy or immaturity which is consequent upon a high development of intelligence, and which must have necessitated the gradual grouping together of pithecoid men into more or less definite families." (See "Descent," I., page 13, on the prolonged infancy of the anthropoid apes.))

Down, November 9th, 1871.

I am greatly obliged to you for having sent me, through my son, your lectures, and for the very honourable manner in which you allude to my works. The lectures seem to me to be written with much force, clearness, and originality. You show also a truly extraordinary amount of knowledge of all that has been published on the subject. The type in many parts is so small that, except to young eyes, it is very difficult to read. Therefore I wish that you would reflect on their separate publication, though so much has been published on the subject that the public may possibly have had enough. I hope that this may be your intention, for I do not think I have ever seen the general argument more forcibly put so as to convert unbelievers.

It has surprised and pleased me to see that you and others have detected the falseness of much of Mr. Mivart's reasoning. I wish I had read your lectures a month or two ago, as I have been preparing a new edition of the "Origin," in which I answer some special points, and I believe I should have found your lectures useful; but my MS. is now in the printer's hands, and I have not strength or time to make any more additions.

P.S.—By an odd coincidence, since the above was written I have received your very obliging letter of October 23rd. I did notice the point to which you refer, and will hereafter reflect more over it. I was indeed on the point of putting in a sentence to somewhat of the same effect in the new edition of the "Origin," in relation to the query—Why have not apes advanced in intellect as much as man? but I omitted it on account of the asserted prolonged infancy of the orang. I am also a little doubtful about the distinction between gregariousness and sociability.

...When you come to England I shall have much pleasure in making your acquaintance; but my health is habitually so weak that I have very small power of conversing with my friends as much as I wish. Let me again thank you for your letter. To believe that I have at all influenced the minds of able men is the greatest satisfaction I am capable of receiving.

LETTER 248. TO E. HACKEL. Down, December 27th, 1871.

I thank you for your very interesting letter, which it has given me much pleasure to receive. I never heard of anything so odd as the Prior in the Holy Catholic Church believing in our ape-like progenitors. I much hope that the Jesuits will not dislodge him.

What a wonderfully active man you are! and I rejoice that you have been so successful in your work on sponges. (248/1. "Die Kalkschwamme: eine Monographie; 3 volumes: Berlin, 1872. H.J. Clark published a paper "On the Spongiae Ciliatae as Infusoria flagellata" in the "Mem. Boston Nat. Hist. Soc." Volume I., Part iii., 1866. See Hackel, op. cit., Volume I., page 24.) Your book with sixty plates will be magnificent. I shall be glad to learn what you think of Clark's view of sponges being flagellate infusorians; some observers in this country believe in him. I am glad you are going fully to consider inheritance, which is an all-important subject for us. I do not know whether you have ever read my chapter on pangenesis. My ideas have been almost universally despised, and I suppose that I was foolish to publish them; yet I must still think that there is some truth in them. Anyhow, they have aided me much in making me clearly understand the facts of inheritance.

I have had bad health this last summer, and during two months was able to do nothing; but I have now almost finished a next edition of the "Origin," which Victor Carus is translating. (248/2. See "Life and Letters," III., page 49.) There is not much new in it, except one chapter in which I have answered, I hope satisfactorily, Mr. Mivart's supposed difficulty on the incipient development of useful structures. I have also given my reasons for quite disbelieving in great and sudden modifications. I am preparing an essay on expression in man and the lower animals. It has little importance, but has interested me. I doubt whether my strength will last for much more serious work. I hope, however, to publish next summer the results of my long-continued experiments on the wonderful advantages derived from crossing. I shall continue to work as long as I can, but it does not much signify when I stop, as there are so many good men fully as capable, perhaps more capable, than myself of carrying on our work; and of these you rank as the first.

With cordial good wishes for your success in all your work and for your happiness.

LETTER 249. TO E. RAY LANKESTER. Down, April 15th [1872].

Very many thanks for your kind consideration. The correspondence was in the "Athenaeum." I got some mathematician to make the calculation, and he blundered and caused me much shame. I send scrap of proofs from last edition of the "Origin," with the calculation corrected. What grand work you did at Naples! I can clearly see that you will some day become our first star in Natural History.

(249/1. Here follows the extract from the "Origin," sixth edition, page 51: "The elephant is reckoned the slowest breeder of all known animals, and I have taken some pains to estimate its probable minimum rate of natural increase. It will be safest to assume that it begins breeding when thirty years old, and goes on breeding till ninety years old, bringing forth six young in the interval, and surviving till one hundred years old; if this be so, after a period of from 740 to 750 years, there would be nearly nineteen million elephants alive, descended from the first pair." In the fifth edition, page 75, the passage runs: "If this be so, at the end of the fifth century, there would be alive fifteen million elephants, descended from the first pair" (see "Athenaeum," June 5, July 3, 17, 24, 1869).)

LETTER 250. TO C. LYELL. Down, May 10th [1872].

I received yesterday morning your present of that work to which I, for one, as well as so many others, owe a debt of gratitude never to be forgotten. I have read with the greatest interest all the special additions; and I wish with all my heart that I had the strength and time to read again every word of the whole book. (250/1. "Principles of Geology," Edition XII., 1875.) I do not agree with all your criticisms on Natural Selection, nor do I suppose that you would expect me to do so. We must be content to differ on several points. I differ must about your difficulty (page 496) (250/2. In Chapter XLIII. Lyell treats of "Man considered with reference to his Origin and Geographical Distribution." He criticizes the view that Natural Selection is capable of bringing about any amount of change provided a series of minute transitional steps can be pointed out. "But in reality," he writes, "it cannot be said that we obtain any insight into the nature of the forces by which a higher grade of organisation or instinct is evolved out of a lower one by becoming acquainted with a series of gradational forms or states, each having a very close affinity with the other."..."It is when there is a change from an inferior being to one of superior grade, from a humbler organism to one endowed with new and more exalted attributes, that we are made to feel that, to explain the difficulty, we must obtain some knowledge of those laws of variation of which Mr. Darwin grants that we are at present profoundly ignorant" (op. cit., pages 496-97).) on a higher grade of organisation being evolved out of lower ones. Is not a very clever man a grade above a very dull one? and would not the accumulation of a large number of slight differences of this kind lead to a great difference in the grade of organisation? And I suppose that you will admit that the difference in the brain of a clever and dull man is not much more wonderful than the difference in the length of the nose of any two men. Of course, there remains the impossibility of explaining at present why one man has a longer nose than another. But it is foolish of me to trouble you with these remarks, which have probably often passed through your mind. The end of this chapter (XLIII.) strikes me as admirably and grandly written. I wish you joy at having completed your gigantic undertaking, and remain, my dear Lyell,

Your ever faithful and now very old pupil, CHARLES DARWIN.

LETTER 251. TO J. TRAHERNE MOGGRIDGE. Sevenoaks, October 9th [1872].

I have just received your note, forwarded to me from my home. I thank you very truly for your intended present, and I am sure that your book will interest me greatly. I am delighted that you have taken up the very difficult and most interesting subject of the habits of insects, on which Englishmen have done so little. How incomparably more valuable are such researches than the mere description of a thousand species! I daresay you have thought of experimenting on the mental powers of the spiders by fixing their trap-doors open in different ways and at different angles, and observing what they will do.

We have been here some days, and intend staying some weeks; for I was quite worn out with work, and cannot be idle at home.

I sincerely hope that your health is not worse.

LETTER 252. TO A. HYATT.

(252/1. The correspondence with Professor Hyatt, of Boston, U.S., originated in the reference to his and Professor Cope's theories of acceleration and retardation, inserted in the sixth edition of the "Origin," page 149.

Mr. Darwin, on receiving from Mr. Hyatt a copy of his "Fossil Cephalopods of the Museum of Comparative Zoology. Embryology," from the "Bull. Mus. Comp. Zool." Harvard, Volume III., 1872, wrote as follows (252/2. Part of this letter was published in "Life and Letters," III., page 154.):—)

October 10th, 1872.

I am very much obliged to you for your kindness in having sent me your valuable memoir on the embryology of the extinct cephalopods. The work must have been one of immense labour, and the results are extremely interesting. Permit me to take this opportunity to express my sincere regret at having committed two grave errors in the last edition of my "Origin of Species," in my allusion to yours and Professor Cope's views on acceleration and retardation of development. I had thought that Professor Cope had preceded you; but I now well remember having formerly read with lively interest, and marked, a paper by you somewhere in my library, on fossil cephalopods, with remarks on the subject. (252/3. The paper seems to be "On the Parallelism between the Different Stages of Life in the Individual and those in the Entire Group of the Molluscous Order Tetrabranchiata," from the "Boston. Soc. Nat. Hist. Mem." I., 1866-69, page 193. On the back of the paper is written, "I cannot avoid thinking this paper fanciful.") It seems also that I have quite misrepresented your joint view; this has vexed me much. I confess that I have never been able to grasp fully what you wish to show, and I presume that this must be owing to some dulness on my part...As the case stands, the law of acceleration and retardation seems to me to be a simple [?] statement of facts; but the statement, if fully established, would no doubt be an important step in our knowledge. But I had better say nothing more on the subject, otherwise I shall perhaps blunder again. I assure you that I regret much that I have fallen into two such grave errors.

LETTER 253. A. HYATT TO CHARLES DARWIN.

(253/1. Mr. Hyatt replied in a long letter, of which only a small part is here given.

Cannstadt bei Stuttgart, November 1872.

The letter with which you have honoured me, bearing the date of October 10th, has just reached here after a voyage to America and back.

I have long had it in mind to write you upon the subject of which you speak, but have been prevented by a very natural feeling of distrust in the worthiness and truth of the views which I had to present.

There is certainly no occasion to apologise for not having quoted my paper. The law of acceleration and retardation of development was therein used to explain the appearance of other phenomena, and might, as it did in nearly all cases, easily escape notice.

My relations with Prof. Cope are of the most friendly character; and although fortunate in publishing a few months ahead, I consider that this gives me no right to claim anything beyond such an amount of participation in the discovery, if it may be so called, as the thoroughness and worth of my work entitles me to...

The collections which I have studied, it will be remembered, are fossils collected without special reference to the very minute subdivisions, such as the subdivisions of the Lower or Middle Lias as made by the German authors, especially Quenstedt and Oppel, but pretty well defined for the larger divisions in which the species are also well defined. The condition of the collections as regards names, etc., was chaotic, localities alone, with some few exceptions, accurate. To put this in order they were first arranged according to their adult characteristics. This proving unsatisfactory, I determined to test thoroughly the theory of evolution by following out the developmental history of each species and placing them within their formations, Middle or Upper Lias, Oolite or so, according to the extent to which they represented each other's characteristics. Thus an adult of simple structure being taken as the starting-point which we will call a, another species which was a in its young stage and became b in the adult was placed above it in the zoological series. By this process I presently found that a, then a b and a b c, c representing the adult stage, were very often found; but that practically after passing these two or three stages it did not often happen that a species was found which was a b c in the young and then became d in the adult. But on the other hand I very frequently found one which, while it was a in the young, skipped the stages b and c and became d while still quite young. Then sometimes, though more rarely, a species would be found belonging to the same series, which would be a in the young and with a very faint and fleeting resemblance to d at a later stage, pass immediately while still quite young to the more advanced characteristics represented by e, and hold these as its specific characteristics until old age destroyed them. This skipping is the highest exemplification, or rather manifestation, of acceleration in development. In alluding to the history of diseases and inheritance of characteristics, you in your "Origin of Species" allude to the ordinary manifestation of acceleration, when you speak of the tendency of diseases or characteristics to appear at younger periods in the life of the child than of its parents. This, according to my observations, is a law, or rather mode, of development, which is applicable to all characteristics, and in this way it is possible to explain why the young of later-occurring animals are like the adult stages of those which preceded them in time. If I am not mistaken you have intimated something of this sort also in your first edition, but I have not been able to find it lately. Of course this is a very normal condition of affairs when a series can be followed in this way, beginning with species a, then going through species a b to a b c, then a b d or a c d, and then a d e or simply a e, as it sometimes comes. Very often the acceleration takes place in two closely connected series, thus:

a—ab—abd—ae—-ad

in which one series goes on very regularly, while another lateral offshoot of a becomes d in the adult. This is an actual case which can be plainly shown with the specimens in hand, and has been verified in the collections here. Retardation is entirely Prof. Cope's idea, but I think also easily traceable. It is the opponent of acceleration, so to speak, or the opposite or negative of that mode of development. Thus series may occur in which, either in size or characteristics, they return to former characteristics; but a better discussion of this point you will find in the little treatise which I send by the same mail as this letter, "On Reversions among the Ammonites."

LETTER 254. TO A. HYATT. Down, December 4th, 1872.

I thank you sincerely for your most interesting letter. You refer much too modestly to your own knowledge and judgment, as you are much better fitted to throw light on your own difficult problems than I am.

It has quite annoyed me that I do not clearly understand yours and Prof. Cope's views (254/1. Prof. Cope's views may be gathered from his "Origin of the Fittest" 1887; in this book (page 41) is reprinted his "Origin of Genera" from the "Proc. Philadelph. Acad. Nat. Soc." 1868, which was published separately by the author in 1869, and which we believe to be his first publication on the subject. In the preface to the "Origin of the Fittest," page vi, he sums up the chief points in the "Origin of Genera" under seven heads, of which the following are the most important:—"First, that development of new characters has been accomplished by an ACCELERATION or RETARDATION in the growth of the parts changed...Second, that of EXACT PARALLELISM between the adult of one individual or set of individuals, and a transitional stage of one or more other individuals. This doctrine is distinct from that of an exact parallelism, which had already been stated by von Baer." The last point is less definitely stated by Hyatt in his letter of December 4th, 1872. "I am thus perpetually led to look upon a series very much as upon an individual, and think that I have found that in many instances these afford parallel changes." See also "Lamarck the Founder of Evolution, by A.S. Packard: New York, 1901.) and the fault lies in some slight degree, I think, with Prof. Cope, who does not write very clearly. I think I now understand the terms "acceleration" and "retardation"; but will you grudge the trouble of telling me, by the aid of the following illustration, whether I do understand rightly? When a fresh-water decapod crustacean is born with an almost mature structure, and therefore does not pass, like other decapods, through the Zoea stage, is this not a case of acceleration? Again, if an imaginary decapod retained, when adult, many Zoea characters, would this not be a case of retardation? If these illustrations are correct, I can perceive why I have been so dull in understanding your views. I looked for something else, being familiar with such cases, and classing them in my own mind as simply due to the obliteration of certain larval or embryonic stages. This obliteration I imagined resulted sometimes entirely from that law of inheritance to which you allude; but that it in many cases was aided by Natural Selection, as I inferred from such cases occurring so frequently in terrestrial and fresh-water members of groups, which retain their several embryonic stages in the sea, as long as fitting conditions are present.

Another cause of my misunderstanding was the assumption that in your series

a—ab—abd—ae,————ad

the differences between the successive species, expressed by the terminal letter, was due to acceleration: now, if I understand rightly, this is not the case; and such characters must have been independently acquired by some means.

The two newest and most interesting points in your letter (and in, as far as I think, your former paper) seem to me to be about senile characteristics in one species appearing in succeeding species during maturity; and secondly about certain degraded characters appearing in the last species of a series. You ask for my opinion: I can only send the conjectured impressions which have occurred to me and which are not worth writing. (It ought to be known whether the senile character appears before or after the period of active reproduction.) I should be inclined to attribute the character in both your cases to the laws of growth and descent, secondarily to Natural Selection. It has been an error on my part, and a misfortune to me, that I did not largely discuss what I mean by laws of growth at an early period in some of my books. I have said something on this head in two new chapters in the last edition of the "Origin." I should be happy to send you a copy of this edition, if you do not possess it and care to have it. A man in extreme old age differs much from a young man, and I presume every one would account for this by failing powers of growth. On the other hand the skulls of some mammals go on altering during maturity into advancing years; as do the horns of the stag, the tail-feathers of some birds, the size of fishes etc.; and all such differences I should attribute simply to the laws of growth, as long as full vigour was retained. Endless other changes of structure in successive species may, I believe, be accounted for by various complex laws of growth. Now, any change of character thus induced with advancing years in the individual might easily be inherited at an earlier age than that at which it first supervened, and thus become characteristic of the mature species; or again, such changes would be apt to follow from variation, independently of inheritance, under proper conditions. Therefore I should expect that characters of this kind would often appear in later-formed species without the aid of Natural Selection, or with its aid if the characters were of any advantage. The longer I live, the more I become convinced how ignorant we are of the extent to which all sorts of structures are serviceable to each species. But that characters supervening during maturity in one species should appear so regularly, as you state to be the case, in succeeding species, seems to me very surprising and inexplicable.

With respect to degradation in species towards the close of a series, I have nothing to say, except that before I arrived at the end of your letter, it occurred to me that the earlier and simpler ammonites must have been well adapted to their conditions, and that when the species were verging towards extinction (owing probably to the presence of some more successful competitors) they would naturally become re-adapted to simpler conditions. Before I had read your final remarks I thought also that unfavourable conditions might cause, through the law of growth, aided perhaps by reversion, degradation of character. No doubt many new laws remain to be discovered. Permit me to add that I have never been so foolish as to imagine that I have succeeded in doing more than to lay down some of the broad outlines of the origin of species.

After long reflection I cannot avoid the conviction that no innate tendency to progressive development exists, as is now held by so many able naturalists, and perhaps by yourself. It is curious how seldom writers define what they mean by progressive development; but this is a point which I have briefly discussed in the "Origin." I earnestly hope that you may visit Hilgendorf's famous deposit. Have you seen Weismann's pamphlet "Einfluss der Isolirung," Leipzig, 1872? He makes splendid use of Hilgendorf's admirable observations. (254/2. Hilgendorf, "Monatsb. K. Akad." Berlin, 1866. For a semi-popular account of Hilgendorf's and Hyatt's work on this subject, see Romanes' "Darwin and after Darwin," I., page 201.) I have no strength to spare, being much out of health; otherwise I would have endeavoured to have made this letter better worth sending. I most sincerely wish you success in your valuable and difficult researches.

I have received, and thank you, for your three pamphlets. As far as I can judge, your views seem very probable; but what a fearfully intricate subject is this of the succession of ammonites. (254/3. See various papers in the publications of the "Boston Soc. Nat. Hist." and in the "Bulletin of the Harvard Museum of Comp. Zoology.")

LETTER 255. A. HYATT TO CHARLES DARWIN. Cannstadt bei Stuttgart, December 8th, 1872.

The quickness and earnestness of your reply to my letter gives me the greatest encouragement, and I am much delighted at the unexpected interest which your questions and comments display. What you say about Prof. Cope's style has been often before said to me, and I have remarked in his writings an unsatisfactory treatment of our common theory. This, I think, perhaps is largely due to the complete absorption of his mind in the contemplation of his subject: this seems to lead him to be careless about the methods in which it may be best explained. He has, however, a more extended knowledge than I have, and has in many ways a more powerful grasp of the subject, and for that very reason, perhaps, is liable to run into extremes. You ask about the skipping of the Zoea stage in fresh-water decapods: is this an illustration of acceleration? It most assuredly is, if acceleration means anything at all. Again, another and more general illustration would be, if, among the marine decapods, a series could be formed in which the Zoea stage became less and less important in the development, and was relegated to younger and younger stages of the development, and finally disappeared in those to which you refer. This is the usual way in which the accelerated mode of development manifests itself; though near the lowest or earliest occurring species it is also to be looked for. Perhaps this to which you allude is an illustration somewhat similar to the one which I have spoken of in my series,

a—ab—abc—ae————ad,

which like "a d" comes from the earliest of a series, though I should think from the entire skipping of the Zoea stage that it must be, like "a e," the result of a long line of ancestors. In fact, the essential point of our theory is, that characteristics are ever inherited by the young at earlier periods than they are assumed in due course of growth by the parents, and that this must eventually lead to the extinction or skipping of these characteristics altogether...

Such considerations as these and the fact that near the heads of series or near the latest members of series, and not at the beginning, were usually found the accelerated types, which skipped lower characteristics and developed very suddenly to a higher and more complex standpoint in structure, led both Cope and [myself] into what may be a great error. I see that it has led you at least into the difficulty of which you very rightly complain, and which, I am sorry to see, has cost you some of your valuable time. We presumed that because characteristics were perpetually inherited at earlier stages, that this very concentration of the developed characteristics made room for the production of differences in the adult descendants of any given pair. Further, that in the room thus made other different characteristics must be produced, and that these would necessarily appear earlier in proportion as the species was more or less accelerated, and be greater or less in the same proportion. Finally, that in the most accelerated, such as "a c" or "a d," the difference would be so great as to constitute distinct genera. Cope and I have differed very much, while he acknowledged the action of the accumulated mode of development only when generic characteristics or greater differences were produced, I saw the same mode of development to be applicable in all cases and to all characteristics, even to diseases. So far the facts bore us out, but when we assumed that the adult differences were the result of the accelerated mode of development, we were perhaps upon rather insecure ground. It is evidently this assumption which has led you to misunderstand the theory. Cope founded his belief, that the adult characteristics were also the result of acceleration, if I rightly remember it, mainly upon the class of facts spoken of above in man where a sudden change into two organs may produce entirely new and unexpected differences in the whole organisation, and upon the changes which acceleration appeared to produce in the development of each succeeding species. Your difficulty in understanding the theory and the observations you have made show me at once what my own difficulties have been, but of these I will not speak at present, as my letter is spinning itself out to a fearful length.

(255/1. After speaking of Cope's comparison of acceleration and retardation in evolution to the force of gravity in physical matters Mr. Hyatt goes on:—)

Now it [acceleration] seems to me to explain less and less the origin of adult progressive characteristics or simply differences, and perhaps now I shall get on faster with my work.

LETTER 256. TO A. HYATT. Down, December 14th [1872].

(256/1. In reply to the above letter (255) from Mr. Hyatt.)

Notwithstanding the kind consideration shown in your last sentence, I must thank you for your interesting and clearly expressed letter. I have directed my publisher to send you a copy of the last edition of the "Origin," and you can, if you like, paste in the "From the Author" on next page. In relation to yours and Professor Cope's view on "acceleration" causing a development of new characters, it would, I think, be well if you were to compare the decapods which pass and do not pass through the Zoea stage, and the one group which does (according to Fritz Muller) pass through to the still earlier Nauplius stages, and see if they present any marked differences. You will, I believe, find that this is not the case. I wish it were, for I have often been perplexed at the omission of embryonic stages as well as the acquirement of peculiar stages appearing to produce no special result in the mature form.

(256/2. The remainder of this letter is missing, and the whole of the last sentence is somewhat uncertainly deciphered. (Note by Mr. Hyatt.))

LETTER 257. TO A. HYATT. Down, February 13th, 1877.

I thank you for your very kind, long, and interesting letter. The case is so wonderful and difficult that I dare not express any opinion on it. Of course, I regret that Hilgendorf has been proved to be so greatly in error (257/1. This refers to a controversy with Sandberger, who had attacked Hilgendorf in the "Verh. der phys.-med. Ges. zu Wurzburg," Bd. V., and in the "Jahrb. der Malakol. Ges." Bd. I., to which Hilgendorf replied in the "Zeitschr. d. Deutschen geolog. Ges." Jahrb. 1877. Hyatt's name occurs in Hilgendorf's pages, but we find no reference to any paper of this date; his well-known paper is in the "Boston. Soc. Nat. Hist." 1880. In a letter to Darwin (May 23rd, 1881) Hyatt regrets that he had no opportunity of a third visit to Steinheim, and goes on: "I should then have done greater justice to Hilgendorf, for whom I have such a high respect."), but it is some selfish comfort to me that I always felt so much misgiving that I never quoted his paper. (257/2. In the fifth edition of the "Origin" (page 362), however, Darwin speaks of the graduated forms of Planorbis multiformis, described by Hilgendorf from certain beds in Switzerland, by which we presume he meant the Steinheim beds in Wurtemberg.) The variability of these shells is quite astonishing, and seems to exceed that of Rubus or Hieracium amongst plants. The result which surprises me most is that the same form should be developed from various and different progenitors. This seems to show how potent are the conditions of life, irrespectively of the variations being in any way beneficial.

The production of a species out of a chaos of varying forms reminds me of Nageli's conclusion, as deduced from the study of Hieracium, that this is the common mode in which species arise. But I still continue to doubt much on this head, and cling to the belief expressed in the first edition of the "Origin," that protean or polymorphic species are those which are now varying in such a manner that the variations are neither advantageous nor disadvantageous. I am glad to hear of the Brunswick deposit, as I feel sure that the careful study of such cases is highly important. I hope that the Smithsonian Institution will publish your memoir.

LETTER 258. TO A. DE CANDOLLE. Down, January 18th [1873].

It was very good of you to give up so much of your time to write to me your last interesting letter. The evidence seems good about the tameness of the alpine butterflies, and the fact seems to me very surprising, for each butterfly can hardly have acquired its experience during its own short life. Will you be so good as to thank M. Humbert for his note, which I have been glad to read. I formerly received from a man, not a naturalist, staying at Cannes a similar account, but doubted about believing it. The case, however, does not answer my query—viz., whether butterflies are attracted by bright colours, independently of the supposed presence of nectar?

I must own that I have great difficulty in believing that any temporary condition of the parents can affect the offspring. If it last long enough to affect the health or structure of the parents, I can quite believe the offspring would be modified. But how mysterious a subject is that of generation! Although my hypothesis of pangenesis has been reviled on all sides, yet I must still look at generation under this point of view; and it makes me very averse to believe in an emotion having any effect on the offspring. Allow me to add one word about blushing and shyness: I intended only to say the habit was primordially acquired by attention to the face, and not that each shy man now attended to his personal appearance.

LETTER 259. TO J.D. HOOKER. Down, June 28th, 1873.

I write a line to wish you good-bye, as I hear you are off on Wednesday, and to thank you for the Dionoea, but I cannot make the little creature grow well. I have this day read Bentham's last address, and must express my admiration of it. (259/1. Presidential address to the Linnean Society, read May 24th, 1873.) Perhaps I ought not to do so, as he fairly crushes me with honour.

I am delighted to see how exactly I agree with him on affinities, and especially on extinct forms as illustrated by his flat-topped tree. (259/2. See page 15 of separate copy: "We should then have the present races represented by the countless branchlets forming the flat-topped summit" of a genealogical tree, in which "all we can do is to map out the summit as it were from a bird's-eye view, and under each cluster, or cluster of clusters, to place as the common trunk an imaginary type of a genus, order, or class according to the depth to which we would go.") My recent work leads me to differ from him on one point—viz., on the separation of the sexes. (259/3. On the question of sexuality, see page 10 of Bentham's address. On the back of Mr. Darwin's copy he has written: "As long as lowest organisms free—sexes separated: as soon as they become attached, to prevent sterility sexes united—reseparated as means of fertilisation, adapted [?] for distant [?] organisms,—in the case of animals by their senses and voluntary movements,—with plants the aid of insects and wind, the latter always existed, and long retained." The two words marked [?] are doubtful. The introduction of freedom or attachedness, as a factor in the problem also occurs in "Cross and Self-fertilisation," page 462. I strongly suspect that sexes were primordially in distinct individuals; then became commonly united in the same individual, and then in a host of animals and some few plants became again separated. Do ask Bentham to send a copy of his address to "Dr. H. Muller, Lippstadt, Prussia," as I am sure it will please him GREATLY.

...When in France write me a line and tell me how you get on, and how Huxley is; but do not do so if you feel idle, and writing bothers you.

LETTER 260. TO R. MELDOLA.

(260/1. This letter, with others from Darwin to Meldola, is published in "Charles Darwin and the Theory of Natural Selection," by E.B. Poulton, pages 199 et seq., London, 1896.)

Southampton, August 13th, 1873.

I am much obliged for your present, which no doubt I shall find at Down on my return home. I am sorry to say that I cannot answer your question; nor do I believe that you could find it anywhere even approximately answered. It is very difficult or impossible to define what is meant by a large variation. Such graduate into monstrosities or generally injurious variations. I do not myself believe that these are often or ever taken advantage of under nature. It is a common occurrence that abrupt and considerable variations are transmitted in an unaltered state, or not at all transmitted, to the offspring, or to some of them. So it is with tailless or hornless animals, and with sudden and great changes of colour in flowers. I wish I could have given you any answer.

LETTER 261. TO E.S. MORSE. [Undated.]

I must have the pleasure of thanking you for your kindness in sending me your essay on the Brachiopoda. (261/1. "The Brachiopoda, a Division of Annelida," "Amer. Assoc. Proc." Volume XIX., page 272, 1870, and "Annals and Mag. Nat. Hist." Volume VI., page 267, 1870.) I have just read it with the greatest interest, and you seem to me (though I am not a competent judge) to make out with remarkable clearness an extremely strong case. What a wonderful change it is to an old naturalist to have to look at these "shells" as "worms"; but, as you truly say, as far as external appearance is concerned, the case is not more wonderful than that of cirripedes. I have also been particularly interested by your remarks on the Geological Record, and on the lower and older forms in each great class not having been probably protected by calcareous valves or a shell.

P.S.—Your woodcut of Lingula is most skilfully introduced to compel one to see its likeness to an annelid.

LETTER 262. TO H. SPENCER.

(262/1. Mr. Spencer's book "The Study of Sociology," 1873, was published in the "Contemporary Review" in instalments between May 1872 and October 1873.)

October 31st [1873].

I am glad to receive to-day an advertisement of your book. I have been wonderfully interested by the articles in the "Contemporary." Those were splendid hits about the Prince of Wales and Gladstone. (262/2. See "The Study of Sociology," page 392. Mr. Gladstone, in protest against some words of Mr. Spencer, had said that the appearance of great men "in great crises of human history" were events so striking "that men would be liable to term them providential in a pre-scientific age." On this Mr. Spencer remarks that "in common with the ancient Greek Mr. Gladstone regards as irreligious any explanation of Nature which dispenses with immediate Divine superintendence." And as an instance of the partnership "between the ideas of natural causation and of providential interference," he instances a case where a prince "gained popularity by outliving certain abnormal changes in his blood," and where "on the occasion of his recovery providential aid and natural causation were unitedly recognised by a thanksgiving to God and a baronetcy to the doctor." The passage on Toryism is on page 395, where Mr. Spencer, with his accustomed tolerance, writes: "The desirable thing is that a growth of ideas and feelings tending to produce modification shall be joined with a continuance of ideas and feelings tending to preserve stability." And from this point of view he concludes it to be very desirable that "one in Mr. Gladstone's position should think as he does." The matter is further discussed in the notes to Chapter XVI., page 423.) I never before read a good defence of Toryism. In one place (but I cannot for the life of me recollect where or what it exactly was) I thought that you would have profited by my principle (i.e. if you do not reject it) given in my "Descent of Man," that new characters which appear late in life are those which are transmitted to the same sex alone. I have advanced some pretty strong evidence, and the principle is of great importance in relation to secondary sexual likenesses. (262/3. This refers to Mr. Spencer's discussion of the evolution of the mental traits characteristic of women. At page 377 he points out the importance of the limitation of heredity by sex in this relation. A striking generalisation on this question is given in the "Descent of Man," Edition I., Volume II., page 285: that when the adult male differs from the adult female, he differs in the same way from the young of both sexes. Can this law be applied in the case in which the adult female possesses characters not possessed by the male: for instance, the high degree of intuitive power of reading the mental states of others and of concealing her own—characters which Mr. Spencer shows to be accounted for by the relations between the husband and wife in a state of savagery. If so, the man should resemble "the young of both sexes" in the absence of these special qualities. This seems to be the case with some masculine characteristics, and childishness of man is not without recognition among women: for instance, by Dolly Winthrop in "Silas Marner," who is content with bread for herself, but bakes cake for children and men, whose "stomichs are made so comical, they want a change—they do, I know, God help 'em.") I have applied it to man and woman, and possibly it was here that I thought that you would have profited by the doctrine. I fear that this note will be almost illegible, but I am very tired.