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More Letters of Charles Darwin - Volume I (of II)
by Charles Darwin
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LETTER 263. G.J. ROMANES TO CHARLES DARWIN.

(263/1. This is, we believe, the first letter addressed by the late Mr. Romanes to Mr. Darwin. It was put away with another on the same subject, and inscribed "Romanes on Abortion, with my answer (very important)." Mr. Darwin's answer given below is printed from his rough draft, which is in places barely decipherable. On the subject of these letters consult Romanes, "Darwin and after Darwin," Volume II., page 99, 1895.)

Dunskaith, Parkhill, Ross-shire, July 10th, 1874.

Knowing that you do not dissuade the more attentive of your readers from communicating directly to yourself any ideas they may have upon subjects connected with your writings, I take the liberty of sending the enclosed copy of a letter, which I have recently addressed to Mr. Herbert Spencer. You will perceive that the subject dealt with is the same as that to which a letter of mine in last week's "Nature" [July 2nd, page 164] refers—viz., "Disuse as a Reducing Cause in Species." In submitting this more detailed exposition of my views to your consideration, I should like to state again what I stated in "Nature" some weeks ago, viz., that in propounding the cessation of selection as a reducing cause, I do not suppose that I am suggesting anything which has not occurred to you already. Not only is this principle embodied in the theory set forth in the article on Rudimentary Organs ("Nature," Volume IX.); but it is more than once hinted at in the "Origin," in the passages where rudimentary organs are said to be more variable than others, because no longer under the restraining influence of Natural Selection. And still more distinctly is this principle recognised in page 120.

Thus, in sending you the enclosed letter, I do not imagine that I am bringing any novel suggestions under your notice. As I see that you have already applied the principle in question to the case of artificially-bred structures, I cannot but infer that you have pondered it in connection with naturally-bred structures. What objection, however, you can have seen to this principle in this latter connection, I am unable to divine; and so I think the best course for me to pursue is the one I adopt—viz., to send you my considerations in full.

In the absence of express information, the most natural inference is that the reason you refuse to entertain the principle in question, is because you show the backward tendency of indiscriminate variability [to be] inadequate to contend with the conservative tendency of long inheritance. The converse of this is expressed in the words "That the struggle between Natural Selection on the one hand, and the tendency to reversion and variability on the other hand, will in the course of time cease; and that the most abnormally developed organs may be made constant, I see no reason to doubt" ("Origin," page 121). Certainly not, if, as I doubt not, the word "constant" is intended to bear a relative signification; but to say that constancy can ever become absolute—i.e., that any term of inheritance could secure to an organ a total immunity from the smallest amount of spontaneous variability—to say this would be unwarrantable. Suppose, for instance, that for some reason or other a further increase in the size of a bat's wing should now suddenly become highly beneficial to that animal: we can scarcely suppose that variations would not be forthcoming for Natural Selection to seize upon (unless the limit of possible size has now been reached, which is an altogether distinct matter). And if we suppose that minute variations on the side of increase are thus even now occasionally taking place, much more is it probable that similar variations on the side of decrease are now taking place—i.e., that if the conservative influence of Natural Selection were removed for a long period of time, more variations would ensue below the present size of bat's wings, than above it. To this it may be added, that when the influence of "speedy selection" is removed, it seems in itself highly probable that the structure would, for this reason, become more variable, for the only reason why it ever ceased to be variable (i.e., after attaining its maximum size), was because of the influence of selection constantly destroying those individuals in which a tendency to vary occurred. When, therefore, this force antagonistic to variability was removed, it seems highly probable that the latter principle would again begin to assert itself, and this in a cumulative manner. Those individuals in which a tendency to vary occurred being no longer cut off, they would have as good a chance of leaving progeny to inherit their fluctuating disposition as would their more inflexible companions.

LETTER 264. TO G.J. ROMANES. July 16th, 1874.

I am much obliged for your kind and long communication, which I have read with great interest, as well as your articles in "Nature." The subject seems to me as important and interesting as it is difficult. I am much out of health, and working very hard on a very different subject, so thus I cannot give your remarks the attention which they deserve. I will, however, keep your letter for some later time, when I may again take up the subject. Your letter makes it clearer to me than it ever was before, how a part or organ which has already begun from any cause to decrease, will go on decreasing through so-called spontaneous variability, with intercrossing; for under such circumstances it is very unlikely that there should be variation in the direction of increase beyond the average size, and no reason why there should not be variations of decrease. I think this expresses your view. I had intended this summer subjecting plants to [illegible] conditions, and observing the effects on variation; but the work would be very laborious, yet I am inclined to think it will be hereafter worth the labour.

LETTER 265. TO T. MEEHAN. Down, October 9th, 1874.

I am glad that you are attending to the colours of dioecious flowers; but it is well to remember that their colours may be as unimportant to them as those of a gall, or, indeed, as the colour of an amethyst or ruby is to these gems. Some thirty years ago I began to investigate the little purple flowers in the centre of the umbels of the carrot. I suppose my memory is wrong, but it tells me that these flowers are female, and I think that I once got a seed from one of them; but my memory may be quite wrong. I hope that you will continue your interesting researches.

LETTER 266. TO G. JAGER. Down, February 3rd, 1875.

I received this morning a copy of your work "Contra Wigand," either from yourself or from your publisher, and I am greatly obliged for it. (266/1. Jager's "In Sachen Darwins insbesondere contra Wigand" (Stuttgart, 1874) is directed against A. Wigand's "Der Darwinismus und die Naturforschung Newtons und Cuviers" (Brunswick, 1874).) I had, however, before bought a copy, and have sent the new one to our best library, that of the Royal Society. As I am a very poor german scholar, I have as yet read only about forty pages; but these have interested me in the highest degree. Your remarks on fixed and variable species deserve the greatest attention; but I am not at present quite convinced that there are such independent of the conditions to which they are subjected. I think you have done great service to the principle of evolution, which we both support, by publishing this work. I am the more glad to read it as I had not time to read Wigand's great and tedious volume.

LETTER 267. TO CHAUNCEY WRIGHT. Down, March 13th, 1875.

I write to-day so that there shall be no delay this time in thanking you for your interesting and long letter received this morning. I am sure that you will excuse brevity when I tell you that I am half-killing myself in trying to get a book ready for the press. (267/1. The MS. of "Insectivorous Plants" was got ready for press in March, 1875. Darwin seems to have been more than usually oppressed by the work.) I quite agree with what you say about advantages of various degrees of importance being co-selected (267/2. Mr. Chauncey Wright wrote (February 24th, 1875): "The inquiry as to which of several real uses is the one through which Natural Selection has acted...has for several years seemed to me a somewhat less important question than it seemed formerly, and still appears to most thinkers on the subject...The uses of the rattling of the rattlesnake as a protection by warning its enemies and as a sexual call are not rival uses; neither are the high-reaching and the far-seeing uses of the giraffe's neck 'rivals.'"), and aided by the effects of use, etc. The subject seems to me well worth further development. I do not think I have anywhere noticed the use of the eyebrows, but have long known that they protected the eyes from sweat. During the voyage of the "Beagle" one of the men ascended a lofty hill during a very hot day. He had small eyebrows, and his eyes became fearfully inflamed from the sweat running into them. The Portuguese inhabitants were familiar with this evil. I think you allude to the transverse furrows on the forehead as a protection against sweat; but remember that these incessantly appear on the foreheads of baboons.

P.S.—I have been greatly pleased by the notices in the "Nation."

LETTER 268. TO A. WEISMANN. Down, May 1st, 1875.

I did not receive your essay for some days after your very kind letter, and I read german so slowly that I have only just finished it. (268/1. "Studien zur Descendenz-Theorie" I. "Ueber den Saison-Dimorphismus," 1875. The fact was previously known that two forms of the genus Vanessa which had been considered to be distinct species are only SEASONAL forms of the same species—one appearing in spring, the other in summer. This remarkable relationship forms the subject of the essay.) Your work has interested me greatly, and your conclusions seem well established. I have long felt much curiosity about season-dimorphism, but never could form any theory on the subject. Undoubtedly your view is very important, as bearing on the general question of variability. When I wrote the "Origin" I could not find any facts which proved the direct action of climate and other external conditions. I long ago thought that the time would soon come when the causes of variation would be fully discussed, and no one has done so much as you in this important subject. The recent evidence of the difference between birds of the same species in the N. and S. United States well shows the power of climate. The two sexes of some few birds are there differently modified by climate, and I have introduced this fact in the last edition of my "Descent of Man." (268/2. "Descent of Man," Edition II. (in one volume), page 423. Allen showed that many species of birds are more strongly coloured in the south of the United States, and that sometimes one sex is more affected than the other. It is this last point that bears on Weismann's remarks (loc. cit., pages 44, 45) on Pieris napi. The males of the alpine-boreal form bryoniae hardly differ from those of the German form (var. vernalis), while the females are strikingly different. Thus the character of secondary sexual differences is determined by climate.) I am, therefore, fully prepared to admit the justness of your criticism on sexual selection of lepidoptera; but considering the display of their beauty, I am not yet inclined to think that I am altogether in error.

What you say about reversion (268/3. For instance, the fact that reversion to the primary winter-form may be produced by the disturbing effect of high temperature (page 7).) being excited by various causes, agrees with what I concluded with respect to the remarkable effects of crossing two breeds: namely, that anything which disturbs the constitution leads to reversion, or, as I put the case under my hypothesis of pangenesis, gives a good chance of latent gemmules developing. Your essay, in my opinion, is an admirable one, and I thank you for the interest which it has afforded me.

P.S. I find that there are several points, which I have forgotten. Mr. Jenner Weir has not published anything more about caterpillars, but I have written to him, asking him whether he has tried any more experiments, and will keep back this letter till I receive his answer. Mr. Riley of the United States supports Mr. Weir, and you will find reference to him and other papers at page 426 of the new and much-corrected edition of my "Descent of Man." As I have a duplicate copy of Volume I. (I believe Volume II. is not yet published in german) I send it to you by this post. Mr. Belt, in his travels in Nicaragua, gives several striking cases of conspicuously coloured animals (but not caterpillars) which are distasteful to birds of prey: he is an excellent observer, and his book, "The Naturalist in Nicaragua," very interesting.

I am very much obliged for your photograph, which I am particularly glad to possess, and I send mine in return.

I see you allude to Hilgendorf's statements, which I was sorry to see disputed by some good German observer. Mr. Hyatt, an excellent palaeontologist of the United States, visited the place, and likewise assured me that Hilgendorf was quite mistaken. (268/4. See Letters 252-7.)

I am grieved to hear that your eyesight still continues bad, but anyhow it has forced your excellent work in your last essay.

May 4th. Here is what Mr. Weir says:—

"In reply to your inquiry of Saturday, I regret that I have little to add to my two communications to the 'Entomological Society Transactions.'

"I repeated the experiments with gaudy caterpillars for years, and always with the same results: not on a single occasion did I find richly coloured, conspicuous larvae eaten by birds. It was more remarkable to observe that the birds paid not the slightest attention to gaudy caterpillars, not even when in motion,—the experiments so thoroughly satisfied my mind that I have now given up making them."

LETTER 269. TO LAWSON TAIT.

(269/1. The late Mr. Lawson Tait wrote to Mr. Darwin (June 2nd, 1875): "I am watching a lot of my mice from whom I removed the tails at birth, and I am coming to the conclusion that the essential use of the tail there is as a recording organ—that is, they record in their memories the corners they turn and the height of the holes they pass through by touching them with their tails." Mr. Darwin was interested in the idea because "some German sneered at Natural Selection and instanced the tails of mice.")

June 11th, 1875.

It has just occurred to me to look at the "Origin of Species" (Edition VI., page 170), and it is certain that Bronn, in the appended chapter to his translation of my book into german, did advance ears and tail of various species of mice as a difficulty opposed to Natural Selection. I answered with respect to ears by alluding to Schobl's curious paper (I forget when published) (269/2. J. Schobl, "Das aussere Ohr der Mause als wichtiges Tastorgan." "Archiv. Mik. Anat." VII., 1871, page 260.) on the hairs of the ears being sensitive and provided with nerves. I presume he made fine sections: if you are accustomed to such histological work, would it not be worth while to examine hairs of tail of mice? At page 189 I quote Henslow (confirmed by Gunther) on Mus messorius (and other species?) using tail as prehensile organ.

Dr. Kane in his account of the second Grinnell Expedition says that the Esquimaux in severe weather carry a fox-tail tied to the neck, which they use as a respirator by holding the tip of the tail between their teeth. (269/3. The fact is stated in Volume II., page 24, of E.K. Kane's "Arctic Explorations: The Second Grinnell Expedition in Search of Sir John Franklin." Philadelphia, 1856.)

He says also that he found a frozen fox curled up with his nose buried in his tail.

N.B. It is just possible that the latter fact is stated by M'Clintock, not by Dr. Kane.

(269/4. The final passage is a postscript by Mr. W.E. Darwin bearing on Mr. Lawson Tait's idea of the respirator function of the fox's tail.)

LETTER 270. TO G.J. ROMANES. Down, July 12th, 1875.

I am correcting a second edition of "Variation under Domestication," and find that I must do it pretty fully. Therefore I give a short abstract of potato graft-hybrids, and I want to know whether I did not send you a reference about beet. Did you look to this, and can you tell me anything about it?

I hope with all my heart that you are getting on pretty well with your experiments.

I have been led to think a good deal on the subject, and am convinced of its high importance, though it will take years of hammering before physiologists will admit that the sexual organs only collect the generative elements.

The edition will be published in November, and then you will see all that I have collected, but I believe that you gave all the more important cases. The case of vine in "Gardeners' Chronicle," which I sent you, I think may only be a bud-variation not due to grafting. I have heard indirectly of your splendid success with nerves of medusae. We have been at Abinger Hall for a month for rest, which I much required, and I saw there the cut-leaved vine which seems splendid for graft hybridism.

LETTER 271. TO FRANCIS GALTON. Down, November 7th, 1875.

I have read your essay with much curiosity and interest, but you probably have no idea how excessively difficult it is to understand. (271/1. "A Theory of Heredity" ("Journal of the Anthropological Institute," 1875). In this paper Mr. Galton admits that the hypothesis of organic units "must lie at the foundation of the science of heredity," and proceeds to show in what respect his conception differs from the hypothesis of pangenesis. The copy of Mr. Galton's paper, which Darwin numbered in correspondence with the criticisms in his letter, is not available, and we are therefore only able to guess at some of the points referred to.) I cannot fully grasp, only here and there conjecture, what are the points on which we differ. I daresay this is chiefly due to muddy-headedness on my part, but I do not think wholly so. Your many terms, not defined, "developed germs," "fertile," and "sterile germs" (the word "germ" itself from association misleading to me) "stirp," "sept," "residue," etc., etc., quite confounded me. If I ask myself how you derive, and where you place the innumerable gemmules contained within the spermatozoa formed by a male animal during its whole life, I cannot answer myself. Unless you can make several parts clearer I believe (though I hope I am altogether wrong) that only a few will endeavour or succeed in fathoming your meaning. I have marked a few passages with numbers, and here make a few remarks and express my opinion, as you desire it, not that I suppose it will be of any use to you.

1. If this implies that many parts are not modified by use and disuse during the life of the individual, I differ widely from you, as every year I come to attribute more and more to such agency. (271/2. This seems to refer to page 329 of Mr. Galton's paper. The passage must have been hastily read, and has been quite misunderstood. Mr. Galton has never expressed the view attributed to him.)

2. This seems rather bold, as sexuality has not been detected in some of the lowest forms, though I daresay it may hereafter be. (271/3. Mr. Galton, op. cit., pages 332-3: "There are not of a necessity two sexes, because swarms of creatures of the simplest organisations mainly multiply by some process of self-division.")

3. If gemmules (to use my own term) were often deficient in buds, I cannot but think that bud-variations would be commoner than they are in a state of nature; nor does it seem that bud-variations often exhibit deficiencies which might be accounted for by the absence of the proper gemmules. I take a very different view of the meaning or cause of sexuality. (271/4. Mr. Galton's idea is that in a bud or other asexually produced part, the germs (i.e. gemmules) may not be completely representative of the whole organism, and if reproduction is continued asexually "at each successive stage there is always a chance of some one or more of the various species of germs... dying out" (page 333). Mr. Galton supposes, in sexual reproduction, where two parents contribute germs to the embryo the chance of deficiency of any of the necessary germs is greatly diminished. Darwin's "very different view of the meaning or cause of sexuality" is no doubt that given in "Cross and Self Fertilisation"—i.e., that sexuality is equivalent to changed conditions, that the parents are not representative of different sexes, but of different conditions of life.)

4. I have ordered "Fraser's Magazine" (271/5. "The History of Twins," by F. Galton, "Fraser's Magazine," November, 1875, republished with additions in the "Journal of the Anthropological Institute," 1875. Mr. Galton explains the striking dissimilarity of twins which is sometimes met with by supposing that the offspring in this case divide the available gemmules between them in such a way that each is the complement of the other. Thus, to put the case in an exaggerated way, similar twins would each have half the gemmules A, B, C,...Z., etc, whereas, in the case of dissimilar twins, one would have all the gemmules A, B, C, D,...M, and the other would have N...Z.), and am curious to learn how twins from a single ovum are distinguished from twins from two ova. Nothing seems to me more curious than the similarity and dissimilarity of twins.

5. Awfully difficult to understand.

6. I have given almost the same notion.

7. I hope that all this will be altered. I have received new and additional cases, so that I have now not a shadow of doubt.

8. Such cases can hardly be spoken of as very rare, as you would say if you had received half the number of cases I have.

(271/6. We are unable to determine to what paragraphs 5, 6, 7, 8 refer.)

I am very sorry to differ so much from you, but I have thought that you would desire my open opinion. Frank is away, otherwise he should have copied my scrawl.

I have got a good stock of pods of sweet peas, but the autumn has been frightfully bad; perhaps we may still get a few more to ripen.

LETTER 272. TO T.H. HUXLEY. Down, November 12th [1875].

Many thanks for your "Biology," which I have read. (272/1. "A Course of Practical Instruction in Elementary Biology," by T.H. Huxley and H.N. Martin, 1875. For an account of the book see "Life and Letters of T.H. Huxley," Volume I., page 380.) It was a real stroke of genius to think of such a plan. Lord, how I wish I had gone through such a course!

LETTER 273. TO FRANCIS GALTON. December 18th [1875].

George has been explaining our differences. I have admitted in the new edition (273/1. In the second edition (1875) of the "Variation of Animals and Plants," Volume II., page 350, reference is made to Mr. Galton's transfusion experiments, "Proc. R. Soc." XIX., page 393; also to Mr. Galton's letter to "Nature," April 27th, 1871, page 502. This is a curious mistake; the letter in "Nature," April 27th, 1871, is by Darwin himself, and refers chiefly to the question whether gemmules may be supposed to be in the blood. Mr. Galton's letter is in "Nature," May 4th, 1871, Volume IV., page 5. See Letter 235.) (before seeing your essay) that perhaps the gemmules are largely multiplied in the reproductive organs; but this does not make me doubt that each unit of the whole system also sends forth its gemmules. You will no doubt have thought of the following objection to your views, and I should like to hear what your answer is. If two plants are crossed, it often, or rather generally, happens that every part of stem, leaf, even to the hairs, and flowers of the hybrid are intermediate in character; and this hybrid will produce by buds millions on millions of other buds all exactly reproducing the intermediate character. I cannot doubt that every unit of the hybrid is hybridised and sends forth hybridised gemmules. Here we have nothing to do with the reproductive organs. There can hardly be a doubt from what we know that the same thing would occur with all those animals which are capable of budding, and some of these (as the compound Ascidians) are sufficiently complex and highly organised.

LETTER 274. TO LAWSON TAIT. March 25th, 1876.

(274/1. The reference is to the theory put forward in the first edition of "Variation of Animals and Plants," II., page 15, that the asserted tendency to regeneration after the amputation of supernumerary digits in man is a return to the recuperative powers characteristic of a "lowly organised progenitor provided with more than five digits." Darwin's recantation is at Volume I., page 459 of the second edition.)

Since reading your first article (274/2. Lawson Tait wrote two notices on "The Variation of Animals and Plants under Domestication" in the "Spectator" of March 4th, 1876, page 312, and March 25th, page 406.), Dr. Rudinger has written to me and sent me an essay, in which he gives the results of the MOST EXTENSIVE inquiries from all eminent surgeons in Germany, and all are unanimous about non-growth of extra digits after amputation. They explain some apparent cases, as Paget did to me. By the way, I struck out of my second edition a quotation from Sir J. Simpson about re-growth in the womb, as Paget demurred, and as I could not say how a rudiment of a limb due to any cause could be distinguished from an imperfect re-growth. Two or three days ago I had another letter from Germany from a good naturalist, Dr. Kollmann (274/3. Dr. Kollmann was Secretary of the Anthropologische Gesellschaft of Munich, in which Society took place the discussion referred to in "Variation of Animals and Plants," I., 459, as originating Darwin's doubts on the whole question. The fresh evidence adduced by Kollmann as to the normal occurrence of a rudimentary sixth digit in Batrachians is Borus' paper, "Die sechste Zehe der Anuren" in "Morpholog. Jahrbuch," Bd. I., page 435. On this subject see Letter 178.), saying he was sorry that I had given up atavism and extra digits, and telling me of new and good evidence of rudiments of a rudimentary sixth digit in Batrachians (which I had myself seen, but given up owing to Gegenbaur's views); but, with re-growth failing me, I could not uphold my old notion.

LETTER 275. TO G.J. ROMANES.

(275/1. Mr. Romanes' reply to this letter is printed in his "Life and Letters," page 93, where by an oversight it is dated 1880-81.)

H. Wedgwood, Esq., Hopedene, Dorking, May 29th [1876].

As you are interested in pangenesis, and will some day, I hope, convert an "airy nothing" into a substantial theory, I send by this post an essay by Hackel (275/2. "Die Perigenesis der Plastidule oder die Wellenzeugung der Lebenstheilchen," 79 pages. Berlin, 1876.) attacking Pan. and substituting a molecular hypothesis. If I understand his views rightly, he would say that with a bird which strengthened its wings by use, the formative protoplasm of the strengthened parts became changed, and its molecular vibrations consequently changed, and that these vibrations are transmitted throughout the whole frame of the bird, and affect the sexual elements in such a manner that the wings of the offspring are developed in a like strengthened manner. I imagine he would say, in cases like those of Lord Morton's mare (275/3. A nearly pure-bred Arabian chestnut mare bore a hybrid to a quagga, and subsequently produced two striped colts by a black Arabian horse: see "Animals and Plants," I., page 403. The case was originally described in the "Philosophical Transactions," 1821, page 20. For an account of recent work bearing on this question, see article on "Zebras, Horses, and Hybrids," in the "Quarterly Review," October 1899. See Letter 235.), that the vibrations from the protoplasm, or "plasson," of the seminal fluid of the zebra set plasson vibrating in the mare; and that these vibrations continued until the hair of the second colt was formed, and which consequently became barred like that of a zebra. How he explains reversion to a remote ancestor, I know not. Perhaps I have misunderstood him, though I have skimmed the whole with some care. He lays much stress on inheritance being a form of unconscious memory, but how far this is part of his molecular vibration, I do not understand. His views make nothing clearer to me; but this may be my fault. No one, I presume, would doubt about molecular movements of some kind. His essay is clever and striking. If you read it (but you must not on my account), I should much like to hear your judgment, and you can return it at any time. The blue lines are Hackel's to call my attention.

We have come here for rest for me, which I have much needed; and shall remain here for about ten days more, and then home to work, which is my sole pleasure in life. I hope your splendid Medusa work and your experiments on pangenesis are going on well. I heard from my son Frank yesterday that he was feverish with a cold, and could not dine with the physiologists, which I am very sorry for, as I should have heard what they think about the new Bill. I see that you are one of the secretaries to this young Society.

LETTER 276. TO H.N. MOSELEY. Down, November 22nd [1876].

It is very kind of you to send me the Japanese books, which are extremely curious and amusing. My son Frank is away, but I am sure he will be much obliged for the two papers which you have sent him.

Thanks, also, for your interesting note. It is a pity that Peripatus (276/1. Moseley "On the Structure and Development of Peripatus capensis" ("Phil. Trans. R. Soc." Volume 164, page 757, 1874). "When suddenly handled or irritated, they (i.e. Peripatus) shoot out fine threads of a remarkably viscid and tenacious milky fluid... projected from the tips of the oral papillae" (page 759).) is so stupid as to spit out the viscid matter at the wrong end of its body; it would have been beautiful thus to have explained the origin of the spider's web.

LETTER 277. NAPHTALI LEWY TO CHARLES DARWIN.

(277/1. The following letter refers to a book, "Toledoth Adam," written by a learned Jew with the object of convincing his co-religionists of the truth of the theory of evolution. The translation we owe to the late Henry Bradshaw, University Librarian at Cambridge. The book is unfortunately no longer to be found in Mr. Darwin's library.)

[1876].

To the Lord, the Prince, who "stands for an ensign of the people" (Isa. xi. 10), the Investigator of the generation, the "bright son of the morning" (Isa. xiv. 12), Charles Darwin, may he live long!

"From the rising of the sun and from the west" (Isa. xlv. 6) all the nations know concerning the Torah (Theory) (277/2. Lit., instruction. The Torah is the Pentateuch, strictly speaking, the source of all knowledge.) which has "proceeded from thee for a light of the people" (Isa. li. 4), and the nations "hear and say, It is truth" (Isa. xliii. 9). But with "the portion of my people" (Jer. x. 16), Jacob, "the lot of my inheritance" (Deut. xxxii. 9), it is not so. This nation, "the ancient people" (Isa. xliv. 7), which "remembers the former things and considers the things of old (Isa. xliii. 18), "knows not, neither doth it understand" (Psalm lxxxii. 5), that by thy Torah (instruction or theory) thou hast thrown light upon their Torah (the Law), and that the eyes of the Hebrews (277/3. One letter in this word changed would make the word "blind," which is what Isaiah uses in the passage alluded to.) "can now see out of obscurity and out of darkness" (Isa. xxix. 18). Therefore "I arose" (Judges v. 7) and wrote this book, "Toledoth Adam" ("the generations of man," Gen. v. 1), to teach the children of my people, the seed of Jacob, the Torah (instruction) which thou hast given for an inheritance to all the nations of the earth.

And I have "proceeded to do a marvellous work among this people, even a marvellous work and a wonder" (Isa. xxix. 14), enabling them now to read in the Torah of Moses our teacher, "plainly and giving the sense" (Neh. viii. 8), that which thou hast given in thy Torahs (works of instruction). And when my people perceive that thy view has by no means "gone astray" (Num. v. 12, 19, etc.) from the Torah of God, they will hold thy name in the highest reverence, and "will at the same time glorify the God of Israel" (Isa. xxix. 23).

"The vision of all this" (Isa. xxix. 11) thou shalt see, O Prince of Wisdom, in this book, "which goeth before me" (Gen. xxxii. 21); and whatever thy large understanding finds to criticise in it, come, "write it in a table and note it in a book" (Isa. xxx. 8); and allow me to name my work with thy name, which is glorified and greatly revered by

Thy servant, Naphtali Hallevi [i.e. the Levite].

Dated here in the city of Radom, in the province of Poland, in the month of Nisan in the year 636, according to the lesser computation (i.e. A.M. [5]636 = A.D. 1876).

LETTER 278. TO OTTO ZACHARIAS. 1877.

When I was on board the "Beagle" I believed in the permanence of species, but, as far as I can remember, vague doubts occasionally flitted across my mind. On my return home in the autumn of 1836 I immediately began to prepare my journal for publication, and then saw how many facts indicated the common descent of species (278/1. "The facts to which reference is here made were, without doubt, eminently fitted to attract the attention of a philosophical thinker; but until the relations of the existing with the extinct species and of the species of the different geographical areas, with one another were determined with some exactness, they afforded but an unsafe foundation for speculation. It was not possible that this determination should have been effected before the return of the "Beagle" to England; and thus the date which Darwin (writing in 1837) assigns to the dawn of the new light which was rising in his mind becomes intelligible."—From "Darwiniana," Essays by Thomas H. Huxley, London, 1893; pages 274-5.), so that in July, 1837, I opened a notebook to record any facts which might bear on the question; but I did not become convinced that species were mutable until, I think, two or three years had elapsed. (278/2. On this last point see page 38.)

LETTER 279. TO G.J. ROMANES.

(279/1. The following letter refers to MS. notes by Romanes, which we have not seen. Darwin's remarks on it are, however, sufficiently clear.)

My address will be "Bassett, Southampton," June 11th [1877].

I have received the crossing paper which you were so kind as to send me. It is very clear, and I quite agree with it; but the point in question has not been a difficulty to me, as I have never believed in a new form originating from a single variation. What I have called unconscious selection by man illustrates, as it seems to me, the same principle as yours, within the same area. Man purchases the individual animals or plants which seem to him the best in any respect—some more so, and some less so—and, without any matching or pairing, the breed in the course of time is surely altered. The absence in numerous instances of intermediate or blending forms, in the border country between two closely allied geographical races or close species, seemed to me a greater difficulty when I discussed the subject in the "Origin."

With respect to your illustration, it formerly drove me half mad to attempt to account for the increase or diminution of the productiveness of an organism; but I cannot call to mind where my difficulty lay. (279/2. See Letters 209-16.) Natural Selection always applies, as I think, to each individual and its offspring, such as its seeds, eggs, which are formed by the mother, and which are protected in various ways. (279/3. It was in regard to this point that Romanes had sent the MS. to Darwin. In a letter of June 16th he writes: "It was with reference to the possibility of Natural Selection acting on organic types as distinguished from individuals,—a possibility which you once told me did not seem at all clear.") There does not seem any difficulty in understanding how the productiveness of an organism might be increased; but it was, as far as I can remember, in reducing productiveness that I was most puzzled. But why I scribble about this I know not.

I have read your review of Mr. Allen's book (279/4. See "Nature" (June 7th, 1877, page 98), a review of Grant Allen's "Physiological Aesthetics."), and it makes me more doubtful, even, than I was before whether he has really thrown much light on the subject.

I am glad to hear that some physiologists take the same view as I did about your giving too much credit to H. Spencer—though, heaven knows, this is a rare fault. (279/5. The reference is to Romanes' lecture on Medusa, given at the Royal Institution, May 25th. (See "Nature," XVI., pages 231, 269, 289.) It appears from a letter of Romanes (June 6th) that it was the abstract in the "Times" that gave the impression referred to. References to Mr. Spencer's theories of nerve-genesis occur in "Nature," pages 232, 271, 289.)

The more I think of your medusa-nerve-work the more splendid it seems to me.

LETTER 280. TO A. DE CANDOLLE. Down, August 3rd, 1877.

I must have the pleasure of thanking you for your long and interesting letter. The cause and means of the transition from an hermaphrodite to a unisexual condition seems to me a very perplexing problem, and I shall be extremely glad to read your remarks on Smilax, whenever I receive the essay which you kindly say that you will send me. (280/1. "Monographiae Phanerogamarum," Volume I. In his treatment of the Smilaceae, De Candolle distinguishes:—Heterosmilax which has dioecious flowers without a trace of aborted stamens or pistils, Smilax with sterile stamens in the female flowers, and Rhipogonum with hermaphrodite flowers.) There is much justice in your criticisms (280/2. The passage criticised by De Candolle is in "Forms of Flowers" (page 7): "It is a natural inference that their corollas have been increased in size for this special purpose." De Candolle goes on to give an account of the "recherche linguistique," which, with characteristic fairness, he undertook to ascertain whether the word "purpose" differs in meaning from the corresponding French word "but.") on my use of the terms object, end, purpose; but those who believe that organs have been gradually modified for Natural Selection for a special purpose may, I think, use the above terms correctly, though no conscious being has intervened. I have found much difficulty in my occasional attempts to avoid these terms, but I might perhaps have always spoken of a beneficial or serviceable effect. My son Francis will be interested by hearing about Smilax. He has dispatched to you a copy of his paper on the glands of Dipsacus (280/3. "Quart. Journ. Mic. Sci." 1877.), and I hope that you will find time to read it, for the case seems to me a new and highly remarkable one. We are now hard at work on an attempt to make out the function or use of the bloom or waxy secretion on the leaves and fruit of many plants; but I doubt greatly whether our experiments will tell us much. (280/4. "As it is we have made out clearly that with some plants (chiefly succulent) the bloom checks evaporation—with some certainly prevents attacks of insects; with some sea-shore plants prevents injury from salt-water, and I believe, with a few prevents injury from pure water resting on the leaves." (See letter to Sir W. Thiselton-Dyer, "Life and Letters," III., page 341. A paper on the same subject by Francis Darwin was published in the "Journ. Linn. Soc." XXII.)) If you have any decided opinion whether plants with conspicuously glaucous leaves are more frequent in hot than in temperate or cold, in dry than in damp countries, I should be grateful if you would add to your many kindnesses by informing me. Pray give my kind remembrances to your son, and tell him that my son has been trying on a large scale the effects of feeding Drosera with meat, and the results are most striking and far more favourable than I anticipated.

LETTER 281. TO G.J. ROMANES.

(281/1. Published in the "Life and Letters" of Romanes, page 66.)

Down, Saturday Night [1877].

I have just finished your lecture (281/2. "The Scientific Evidence of Organic Evolution: a Discourse" (delivered before the Philosophical Society of Ross-shire), Inverness, 1877. It was reprinted in the "Fortnightly Review," and was afterwards worked up into a book under the above title.); it is an admirable scientific argument, and most powerful. I wish that it could be sown broadcast throughout the land. Your courage is marvellous, and I wonder that you were not stoned on the spot—and in Scotland! Do please tell me how it was received in the Lecture Hall. About man being made like a monkey (page 37 (281/3. "And if you reject the natural explanation of hereditary descent, you can only suppose that the Deity, in creating man, took the most scrupulous pains to make him in the image of the ape" ("Discourse," page 37).)) is quite new to me, and the argument in an earlier place (page 8 (281/4. At page 8 of the "Discourse" the speaker referred to the law "which Sir William Hamilton called the Law of Parsimony—or the law which forbids us to assume the operation of higher causes when lower ones are found sufficient to explain the desired effects," as constituting the "only logical barrier between Science and Superstition.")) on the law of parsimony admirably put. Yes, page 21 (281/5. "Discourse," page 21. If we accept the doctrines of individual creations and ideal types, we must believe that the Deity acted "with no other apparent motive than to suggest to us, by every one of the observable facts, that the ideal types are nothing other than the bonds of a lineal descent.") is new to me. All strike me as very clear, and, considering small space, you have chosen your lines of reasoning excellently.

The few last pages are awfully powerful, in my opinion.

Sunday Morning.—The above was written last night in the enthusiasm of the moment, and now—this dark, dismal Sunday morning—I fully agree with what I said.

I am very sorry to hear about the failures in the graft experiments, and not from your own fault or ill-luck. Trollope in one of his novels gives as a maxim of constant use by a brickmaker—"It is dogged as does it" (281/6. "Tell 'ee what, Master Crawley;—and yer reverence mustn't think as I means to be preaching; there ain't nowt a man can't bear if he'll only be dogged. You go whome, Master Crawley, and think o' that, and may be it'll do ye a good yet. It's dogged as does it. It ain't thinking about it." (Giles Hoggett, the old Brickmaker, in "The Last Chronicle of Barset," Volume II., 1867, page 188.))—and I have often and often thought that this is the motto for every scientific worker. I am sure it is yours—if you do not give up pangenesis with wicked imprecations.

By the way, G. Jager has brought out in "Kosmos" a chemical sort of pangenesis bearing chiefly on inheritance. (281/7. Several papers by Jager on "Inheritance" were published in the first volume of "Kosmos," 1877.)

I cannot conceive why I have not offered my garden for your experiments. I would attend to the plants, as far as mere care goes, with pleasure; but Down is an awkward place to reach.

Would it be worth while to try if the "Fortnightly" would republish it [i.e. the lecture]?

LETTER 282. TO T.H. HUXLEY.

(282/1. In 1877 the honorary degree of LL.D. was conferred on Mr. Darwin by the University of Cambridge. At the dinner given on the occasion by the Philosophical Society, Mr. Huxley responded to the toast of the evening with the speech of which an authorised version is given by Mr. L. Huxley in the "Life and Letters" of his father (Volume I., page 479). Mr. Huxley said, "But whether the that doctrine [of evolution] be true or whether it be false, I wish to express the deliberate opinion, that from Aristotle's great summary of the biological knowledge of his time down to the present day, there is nothing comparable to the "Origin of Species," as a connected survey of the phenomena of life permeated and vivified by a central idea."

In the first part of the speech there was a brilliant sentence which he described as a touch of the whip "tied round with ribbons," and this was perhaps a little hard on the supporters of evolution in the University. Mr. Huxley said "Instead of offering her honours when they ran a chance of being crushed beneath the accumulated marks of approbation of the whole civilised world, the University has waited until the trophy was finished, and has crowned the edifice with the delicate wreath of academic appreciation.")

Down, Monday night, November 19th [1877].

I cannot rest easy without telling you more gravely than I did when we met for five minutes near the Museum, how deeply I have felt the many generous things (as far as Frank could remember them) which you said about me at the dinner. Frank came early next morning boiling over with enthusiasm about your speech. You have indeed always been to me a most generous friend, but I know, alas, too well how greatly you overestimate me. Forgive me for bothering you with these few lines.

(282/2. The following extract from a letter (February 10th, 1878) to his old schoolfellow, Mr. J. Price, gives a characteristic remark about the honorary degree.)

"I am very much obliged for your kind congratulations about the LL.D. Why the Senate conferred it on me I know not in the least. I was astonished to hear that the R. Prof. of Divinity and several other great Dons attended, and several such men have subscribed, as I am informed, for the picture for the University to commemorate the honour conferred on me."

LETTER 283. TO W. BOWMAN.

(283/1. We have not discovered to what prize the following letter to the late Sir W. Bowman (the well known surgeon) refers.)

Down, February 22nd, 1878.

I received your letter this morning, and it was quite impossible that you should receive an answer by 4 p.m. to-day. But this does not signify in the least, for your proposal seems to me a very good one, and I most entirely agree with you that it is far better to suggest some special question rather than to have a general discussion compiled from books. The rule that the Essay must be "illustrative of the wisdom and beneficence of the Almighty" would confine the subjects to be proposed. With respect to the Vegetable Kingdom, I could suggest two or three subjects about which, as it seems to me, information is much required; but these subjects would require a long course of experiment, and unfortunately there is hardly any one in this country who seems inclined to devote himself to experiments.

LETTER 284. TO J. TORBITT.

(284/1. Mr. Torbitt was engaged in trying to produce by methodical selection and cross-fertilisation a fungus-proof race of the potato. The plan is fully described in the "Life and Letters," III., page 348. The following letter is given in additional illustration of the keen interest Mr. Darwin took in the project.)

Down, Monday, March 4th, 1878.

I have nothing good to report. Mr. Caird called upon me yesterday; both he and Mr. Farrer (284/2. The late Lord Farrer.) have been most energetic and obliging. There is no use in thinking about the Agricultural Society. Mr. Caird has seen several persons on the subject, especially Mr. Carruthers, Botanist to the Society. He (Mr. Carruthers) thinks the attempt hopeless, but advances in a long memorandum sent to Mr. Caird, reasons which I am convinced are not sound. He specifies two points, however, which are well worthy of your consideration—namely, that a variety should be tested three years before its soundness can be trusted; and especially it should be grown under a damp climate. Mr. Carruthers' opinion on this head is valuable because he was employed by the Society in judging the varieties sent in for the prize offered a year or two ago. If I had strength to get up a memorial to Government, I believe that I could succeed; for Sir J. Hooker writes that he believes you are on the right path; but I do not know to whom else to apply whose judgment would have weight with Government, and I really have not strength to discuss the matter and convert persons.

At Mr. Farrer's request, when we hoped the Agricultural Society might undertake it, I wrote to him a long letter giving him my opinion on the subject; and this letter Mr. Caird took with him yesterday, and will consider with Mr. Farrer whether any application can be made to Government.

I am, however, far from sanguine. I shall see Mr. Farrer this evening, and will do what I can. When I receive back my letter I will send it to you for your perusal.

After much reflection it seems to me that your best plan will be, if we fail to get Government aid, to go on during the present year, on a reduced scale, in raising new cross-fertilised varieties, and next year, if you are able, testing the power of endurance of only the most promising kind. If it were possible it would be very advisable for you to get some grown on the wet western side of Ireland. If you succeed in procuring a fungus-proof variety you may rely on it that its merits would soon become known locally and it would afterwards spread rapidly far and wide. Mr. Caird gave me a striking instance of such a case in Scotland. I return home to-morrow morning.

I have the pleasure to enclose a cheque for 100 pounds. If you receive a Government grant, I ought to be repaid.

P.S. If I were in your place I would not expend any labour or money in publishing what you have already done, or in sending seeds or tubers to any one. I would work quietly on till some sure results were obtained. And these would be so valuable that your work in this case would soon be known. I would also endeavour to pass as severe a judgment as possible on the state of the tubers and plants.

LETTER 285. TO E. VON MOJSISOVICS. Down, June 1st, 1878.

I have at last found time to read [the] first chapter of your "Dolomit Riffe" (285/1. "Dolomitriffe Sudtirols und Venetiens." Wien, 1878.), and have been exceedingly interested by it. What a wonderful change in the future of geological chronology you indicate, by assuming the descent-theory to be established, and then taking the graduated changes of the same group of organisms as the true standard! I never hoped to live to see such a step even proposed by any one. (285/2. Published in "Life and Letters," III., pages 234, 235.)

Nevertheless, I saw dimly that each bed in a formation could contain only the organisms proper to a certain depth, and to other there existing conditions, and that all the intermediate forms between one marine species and another could rarely be preserved in the same place and bed. Oppel, Neumayr, and yourself will confer a lasting and admirable service on the noble science of Geology, if you can spread your views so as to be generally known and accepted.

With respect to the continental and oceanic periods common to the whole northern hemisphere, to which you refer, I have sometimes speculated that the present distribution of the land and sea over the world may have formerly been very different to what it now is; and that new genera and families may have been developed on the shores of isolated tracts in the south, and afterwards spread to the north.

LETTER 286. TO J.W. JUDD. Down, June 27th, 1878.

I am heartily glad to hear of your intended marriage. A good wife is the supreme blessing in this life, and I hope and believe from what you say that you will be as happy as I have been in this respect. May your future geological work be as valuable as that which you have already done; and more than this need not be wished for any man. The practical teaching of Geology seems an excellent idea.

Many thanks for Neumayr, (286/1. Probably a paper on "Die Congerien und Paludinenschichten Slavoniens und deren Fauna. Ein Beitrag zur Descendenz-Theorie," "Wien. Geol. Abhandl." VII. (Heft 3), 1874-82.), but I have already received and read a copy of the same, or at least of a very similar essay, and admirably good it seemed to me.

This essay, and one by Mojsisovics (286/2. See note to Letter 285.), which I have lately read, show what Palaeontology in the future will do for the classification and sequence of formations. It delighted me to see so inverted an order of proceeding—viz., the assuming the descent of species as certain, and then taking the changes of closely allied forms as the standard of geological time. My health is better than it was a few years ago, but I never pass a day without much discomfort and the sense of extreme fatigue.

(286/3. We owe to Professor Judd the following interesting recollections of Mr. Darwin, written about 1883:—

"On this last occasion, when I congratulated him on his seeming better condition of health, he told me of the cause for anxiety which he had in the state of his heart. Indeed, I cannot help feeling that he had a kind of presentiment that his end was approaching. When I left him, he insisted on conducting me to the door, and there was that in his tone and manner which seemed to convey to me the sad intelligence that it was not merely a temporary farewell, though he himself was perfectly cheerful and happy.

"It is impossible for me adequately to express the impression made upon my mind by my various conversations with Mr. Darwin. His extreme modesty led him to form the lowest estimate of his own labours, and a correspondingly extravagant idea of the value of the work done by others. His deference to the arguments and suggestions of men greatly his juniors, and his unaffected sympathy in their pursuits, was most marked and characteristic; indeed, he, the great master of science, used to speak, and I am sure felt, as though he were appealing to superior authority for information in all his conversations. It was only when a question was fully discussed with him that one became conscious of the fund of information he could bring to its elucidation, and the breadth of thought with which he had grasped it. Of his gentle, loving nature, of which I had so many proofs, I need not write; no one could be with him, even for a few minutes, without being deeply impressed by his grateful kindliness and goodness.")

LETTER 287. TO COUNT SAPORTA. Down, August 15th, 1878.

I thank you very sincerely for your kind and interesting letter. It would be false in me to pretend that I care very much about my election to the Institute, but the sympathy of some few of my friends has gratified me deeply.

I am extremely glad to hear that you are going to publish a work on the more ancient fossil plants; and I thank you beforehand for the volume which you kindly say that you will send me. I earnestly hope that you will give, at least incidentally, the results at which you have arrived with respect to the more recent Tertiary plants; for the close gradation of such forms seems to me a fact of paramount importance for the principle of evolution. Your cases are like those on the gradation in the genus Equus, recently discovered by Marsh in North America.

LETTER 288. TO THE DUKE OF ARGYLL.

(288/1. The following letter was published in "Nature," March 5th, 1891, Volume XLIII., page 415, together with a note from the late Duke of Argyll, in which he stated that the letter had been written to him by Mr. Darwin in reply to the question, "why it was that he did assume the unity of mankind as descended from a single pair." The Duke added that in the reply Mr. Darwin "does not repudiate this interpretation of his theory, but simply proceeds to explain and to defend the doctrine." On a former occasion the Duke of Argyll had "alluded as a fact to the circumstance that Charles Darwin assumed mankind to have arisen at one place, and therefore in a single pair." The letter from Darwin was published in answer to some scientific friends, who doubted the fact and asked for the reference on which the statement was based.)

Down, September 23rd, 1878.

The problem which you state so clearly is a very interesting one, on which I have often speculated. As far as I can judge, the improbability is extreme that the same well-characterised species should be produced in two distinct countries, or at two distinct times. It is certain that the same variation may arise in two distinct places, as with albinism or with the nectarine on peach-trees. But the evidence seems to me overwhelming that a well-marked species is the product, not of a single or of a few variations, but of a long series of modifications, each modification resulting chiefly from adaptation to infinitely complex conditions (including the inhabitants of the same country), with more or less inheritance of all the preceding modifications. Moreover, as variability depends more on the nature of the organism than on that of the environment, the variations will tend to differ at each successive stage of descent. Now it seems to me improbable in the highest degree that a species should ever have been exposed in two places to infinitely complex relations of exactly the same nature during a long series of modifications. An illustration will perhaps make what I have said clearer, though it applies only to the less important factors of inheritance and variability, and not to adaptation—viz., the improbability of two men being born in two countries identical in body and mind. If, however, it be assumed that a species at each successive stage of its modification was surrounded in two distinct countries or times, by exactly the same assemblage of plants and animals, and by the same physical conditions, then I can see no theoretical difficulty [in] such a species giving birth to the new form in the two countries. If you will look to the sixth edition of my "Origin," at page 100, you will find a somewhat analogous discussion, perhaps more intelligible than this letter.

LETTER 289. W.T. THISELTON-DYER TO THE EDITOR OF "NATURE."

(289/1. The following letter ("Nature," Volume XLIII., page 535) criticises the interpretation given by the Duke to Mr. Darwin's letter.)

Royal Gardens, Kew, March 27th [1891].

In "Nature" of March 5th (page 415), the Duke of Argyll has printed a very interesting letter of Mr. Darwin's, from which he drew the inference that the writer "assumed mankind to have arisen...in a single pair." I do not think myself that the letter bears this interpretation. But the point in its most general aspect is a very important one, and is often found to present some difficulty to students of Mr. Darwin's writings.

Quite recently I have found by accident, amongst the papers of the late Mr. Bentham at Kew, a letter of friendly criticism from Mr. Darwin upon the presidential address which Mr. Bentham delivered to the Linnean Society on May 24th, 1869. This letter, I think, has been overlooked and not published previously. In it Mr. Darwin expresses himself with regard to the multiple origin of races and some other points in very explicit language. Prof. Meldola, to whom I mentioned in conversation the existence of the letter, urged me strongly to print it. This, therefore, I now do, with the addition of a few explanatory notes.

LETTER 290. TO G. BENTHAM. Down, November 25th, 1869.

(290/1. The notes to this letter are by Sir W. Thiselton-Dyer, and appeared in "Nature," loc. cit.)

I was greatly interested by your address, which I have now read thrice, and which I believe will have much influence on all who read it. But you are mistaken in thinking that I ever said you were wrong on any point. All that I meant was that on certain points, and these very doubtful points, I was inclined to differ from you. And now, on further considering the point on which some two or three months ago I felt most inclined to differ—viz., on isolation—I find I differ very little. What I have to say is really not worth saying, but as I should be very sorry not to do whatever you asked, I will scribble down the slightly dissentient thoughts which have occurred to me. It would be an endless job to specify the points in which you have interested me; but I may just mention the relation of the extreme western flora of Europe (some such very vague thoughts have crossed my mind, relating to the Glacial period) with South Africa, and your remarks on the contrast of passive and active distribution.

Page lxx.—I think the contingency of a rising island, not as yet fully stocked with plants, ought always to be kept in mind when speaking of colonisation.

Page lxxiv.—I have met with nothing which makes me in the least doubt that large genera present a greater number of varieties relatively to their size than do small genera. (290/2. Bentham thought "degree of variability... like other constitutional characters, in the first place an individual one, which...may become more or less hereditary, and therefore specific; and thence, but in a very faint degree, generic." He seems to mean to argue against the conclusion which Sir Joseph Hooker had quoted from Mr. Darwin that "species of large genera are more variable than those of small." [On large genera varying, see Letter 53.]) Hooker was convinced by my data, never as yet published in full, only abstracted in the "Origin."

Page lxxviii.—I dispute whether a new race or species is necessarily, or even generally, descended from a single or pair of parents. The whole body of individuals, I believe, become altered together—like our race-horses, and like all domestic breeds which are changed through "unconscious selection" by man. (290/3. Bentham had said: "We must also admit that every race has probably been the offspring of one parent or pair of parents, and consequently originated in one spot." The Duke of Argyll inverts the proposition.)

When such great lengths of time are considered as are necessary to change a specific form, I greatly doubt whether more or less rapid powers of multiplication have more than the most insignificant weight. These powers, I think, are related to greater or less destruction in early life.

Page lxxix.—I still think you rather underrate the importance of isolation. I have come to think it very important from various grounds; the anomalous and quasi-extinct forms on islands, etc., etc., etc.

With respect to areas with numerous "individually durable" forms, can it be said that they generally present a "broken" surface with "impassable barriers"? This, no doubt, is true in certain cases, as Teneriffe. But does this hold with South-West Australia or the Cape? I much doubt. I have been accustomed to look at the cause of so many forms as being partly an arid or dry climate (as De Candolle insists) which indirectly leads to diversified [?] conditions; and, secondly, to isolation from the rest of the world during a very long period, so that other more dominant forms have not entered, and there has been ample time for much specification and adaptation of character.

Page lxxx.—I suppose you think that the Restiaceae, Proteaceae (290/4. It is doubtful whether Bentham did think so. In his 1870 address he says: "I cannot resist the opinion that all presumptive evidence is against European Proteaceae, and that all direct evidence in their favour has broken down upon cross-examination."), etc., etc., once extended over the world, leaving fragments in the south.

You in several places speak of distribution of plants as if exclusively governed by soil and climate. I know that you do not mean this, but I regret whenever a chance is omitted of pointing out that the struggle with other plants (and hostile animals) is far more important.

I told you that I had nothing worth saying, but I have given you my THOUGHTS.

How detestable are the Roman numerals! why should not the President's addresses, which are often, and I am sure in this case, worth more than all the rest of the number, be paged with Christian figures?

LETTER 291. TO R. MELDOLA.

(291/1. "This letter was in reply to a suggestion that in his preface Mr. Darwin should point out by references to "The Origin of Species" and his other writings how far he had already traced out the path which Weismann went over. The suggestion was made because in a great many of the continental writings upon the theory of descent, many of the points which had been clearly foreshadowed, and in some cases even explicitly stated by Darwin, had been rediscovered and published as though original. In the notes to my edition of Weismann I have endeavoured to do Darwin full justice.—R.M." See Letter 310.)

4, Bryanston Street, November 26th, 1878.

I am very sorry to say that I cannot agree to your suggestion. An author is never a fit judge of his own work, and I should dislike extremely pointing out when and how Weismann's conclusions and work agreed with my own. I feel sure that I ought not to do this, and it would be to me an intolerable task. Nor does it seem to me the proper office of the preface, which is to show what the book contains, and that the contents appear to me valuable. But I can see no objection for you, if you think fit, to write an introduction with remarks or criticisms of any kind. Of course, I would be glad to advise you on any point as far as lay in my power, but as a whole I could have nothing to do with it, on the grounds above specified, that an author cannot and ought not to attempt to judge his own works, or compare them with others. I am sorry to refuse to do anything which you wish.

LETTER 292. TO T.H. HUXLEY. Down, January 18th, 1879.

I have just finished your present of the Life of Hume (292/1. "Hume" in Mr. Morley's "English Men of Letters" series. Of the biographical part of this book Mr. Huxley wrote, in a letter to Mr. Skelton, January 1879 ("Life of T.H. Huxley," II., page 7): "It is the nearest approach to a work of fiction of which I have yet been guilty."), and must thank you for the great pleasure which it has given me. Your discussions are, as it seems to me, clear to a quite marvellous degree, and many of the little interspersed flashes of wit are delightful. I particularly enjoyed the pithy judgment in about five words on Comte. (292/2. Possibly the passage referred to is on page 52.) Notwithstanding the clearness of every sentence, the subjects are in part so difficult that I found them stiff reading. I fear, therefore, that it will be too stiff for the general public; but I heartily hope that this will prove to be a mistake, and in this case the intelligence of the public will be greatly exalted in my eyes. The writing of this book must have been awfully hard work, I should think.

LETTER 293. TO F. MULLER. Down, March 4th [1879].

I thank you cordially for your letter. Your facts and discussion on the loss of the hairs on the legs of the caddis-flies seem to me the most important and interesting thing which I have read for a very long time. I hope that you will not disapprove, but I have sent your letter to "Nature" (293/1. Fritz Muller, "On a Frog having Eggs on its Back—On the Abortion of the Hairs on the Legs of certain Caddis-Flies, etc.": Muller's letter and one from Charles Darwin were published in "Nature," Volume XIX., page 462, 1879.), with a few prefatory remarks, pointing out to the general reader the importance of your view, and stating that I have been puzzled for many years on this very point. If, as I am inclined to believe, your view can be widely extended, it will be a capital gain to the doctrine of evolution. I see by your various papers that you are working away energetically, and, wherever you look, you seem to discover something quite new and extremely interesting. Your brother also continues to do fine work on the fertilisation of flowers and allied subjects.

I have little or nothing to tell you about myself. I go slowly crawling on with my present subject—the various and complicated movements of plants. I have not been very well of late, and am tired to-day, so will write no more. With the most cordial sympathy in all your work, etc.

LETTER 294. TO T.H. HUXLEY. Down, April 19th, 1879.

Many thanks for the book. (294/1. Ernst Hackel's "Freedom in Science and Teaching," with a prefatory note by T.H. Huxley, 1879. Professor Hackel has recently published (without permission) a letter in which Mr. Darwin comments severely on Virchow. It is difficult to say which would have pained Mr. Darwin more—the affront to a colleague, or the breach of confidence in a friend.) I have read only the preface...It is capital, and I enjoyed the tremendous rap on the knuckles which you gave Virchow at the close. What a pleasure it must be to write as you can do!

LETTER 295. TO E.S. MORSE. Down, October 21st, 1879.

Although you are so kind as to tell me not to write, I must just thank you for the proofs of your paper, which has interested me greatly. (295/1. See "The Shell Mounds of Omori" in the "Memoirs of the Science Department of the Univ. of Tokio," Volume I., Part I., 1879. The ridges on Arca are mentioned at page 25. In "Nature," April 15th, 1880, Mr. Darwin published a letter by Mr. Morse relating to the review of the above paper, which appeared in "Nature," XXI., page 350. Mr. Darwin introduces Mr. Morse's letter with some prefatory remarks. The correspondence is republished in the "American Naturalist," September, 1880.) The increase in the number of ridges in the three species of Arca seems to be a very noteworthy fact, as does the increase of size in so many, yet not all, the species. What a constant state of fluctuation the whole organic world seems to be in! It is interesting to hear that everywhere the first change apparently is in the proportional numbers of the species. I was much struck with the fact in the upraised shells of Coquimbo, in Chili, as mentioned in my "Geological Observations on South America."

Of all the wonders in the world, the progress of Japan, in which you have been aiding, seems to me about the most wonderful.

LETTER 296. TO A.R. WALLACE. Down, January 5th 1880.

As this note requires no sort of answer, you must allow me to express my lively admiration of your paper in the "Nineteenth Century." (296/1. "Nineteenth Century," January 1880, page 93, "On the Origin of Species and Genera.") You certainly are a master in the difficult art of clear exposition. It is impossible to urge too often that the selection from a single varying individual or of a single varying organ will not suffice. You have worked in capitally Allen's admirable researches. (296/2. J.A. Allen, "On the Mammals and Winter Birds of East Florida, etc." ("Bull. Mus. Comp. Zoolog. Harvard," Volume II.) As usual, you delight to honour me more than I deserve. When I have written about the extreme slowness of Natural Selection (296/3. Mr. Wallace makes a calculation based on Allen's results as to the very short period in which the formation of a race of birds differing 10 to 20 per cent. from the average in length of wing and strength of beak might conceivably be effected. He thinks that the slowness of the action of Natural Selection really depends on the slowness of the changes naturally occurring in the physical conditions, etc.) (in which I hope I may be wrong), I have chiefly had in my mind the effects of intercrossing. I subscribe to almost everything you say excepting the last short sentence. (296/4. The passage in question is as follows: "I have also attempted to show that the causes which have produced the separate species of one genus, of one family, or perhaps of one order, from a common ancestor, are not necessarily the same as those which have produced the separate orders, classes, and sub-kingdoms from more remote common ancestors. That all have been alike produced by 'descent with modification' from a few primitive types, the whole body of evidence clearly indicates; but while individual variation with Natural Selection is proved to be adequate for the production of the former, we have no proof and hardly any evidence that it is adequate to initiate those important divergences of type which characterise the latter." In this passage stress should be laid (as Mr. Wallace points out to us) on the word PROOF. He by no means asserts that the causes which have produced the species of a genus are inadequate to produce greater differences. His object is rather to urge the difference between proof and probability.)

LETTER 297. TO J.H. FABRE.

(297/1. A letter to M. Fabre is given in "Life and Letters," III., page 220, in which the suggestion is made of rotating the insect before a "homing" experiment occurs.)

Down, February 20th, 1880.

I thank you for your kind letter, and am delighted that you will try the experiment of rotation. It is very curious that such a belief should be held about cats in your country (297/2. M. Fabre had written from Serignan, Vaucluse: "Parmi la population des paysans de mon village, l'habitude est de faire tourner dans un sac le chat que l'on se propose de porter ailleurs, et dont on veut empecher le retour. J'ignore si cette pratique obtient du succes."), I never heard of anything of the kind in England. I was led, as I believe, to think of the experiment from having read in Wrangel's "Travels in Siberia" (297/3. Admiral Ferdinand Petrovich von Wrangell, "Le Nord de la Siberie, Voyage parmi les Peuplades de la Russie asiatique, etc." Paris, 1843.) of the wonderful power which the Samoyedes possess of keeping their direction in a fog whilst travelling in a tortuous line through broken ice. With respect to cats, I have seen an account that in Belgium there is a society which gives prizes to the cat which can soonest find its way home, and for this purpose they are carried to distant parts of the city.

Here would be a capital opportunity for trying rotation.

I am extremely glad to hear that your book will probably be translated into English.

P.S.—I shall be much pleased to hear the result of your experiments.

LETTER 298. TO J.H. FABRE. Down, January 21st, 1881.

I am much obliged for your very interesting letter. Your results appear to me highly important, as they eliminate one means by which animals might perhaps recognise direction; and this, from what has been said about savages, and from our own consciousness, seemed the most probable means. If you think it worth while, you can of course mention my name in relation to this subject.

Should you succeed in eliminating a sense of the magnetic currents of the earth, you would leave the field of investigation quite open. I suppose that even those who still believe that each species was separately created would admit that certain animals possess some sense by which they perceive direction, and which they use instinctively. On mentioning the subject to my son George, who is a mathematician and knows something about magnetism, he suggested making a very thin needle into a magnet; then breaking it into very short pieces, which would still be magnetic, and fastening one of these pieces with some cement on the thorax of the insect to be experimented on.

He believes that such a little magnet, from its close proximity to the nervous system of the insect, would affect it more than would the terrestrial currents.

I have received your essay on Halictus (298/1. "Sur les Moeurs et la Parthenogese des Halictes" ("Ann. Sc. Nat." IX., 1879-80).), which I am sure that I shall read with much interest.

LETTER 299. TO T.H. HUXLEY.

(299/1. On April 9th, 1880, Mr. Huxley lectured at the Royal Institution on "The Coming of Age of the Origin of Species." The lecture was published in "Nature" and in Huxley's "Collected Essays," Volume II., page 227. Darwin's letter to Huxley on the subject is given in "Life and Letters," III., page 240; in Huxley's reply of May 10th ("Life and Letters of T.H. Huxley," II., page 12) he writes: "I hope you do not imagine because I had nothing to say about 'Natural Selection' that I am at all weak of faith on that article...But the first thing seems to me to be to drive the fact of evolution into people's heads; when that is once safe, the rest will come easy.")

Down, May 11th, 1880.

I had no intention to make you write to me, or expectation of your doing so; but your note has been so far "cheerier" (299/2. "You are the cheeriest letter-writer I know": Huxley to Darwin. See Huxley's "Life," II., page 12.) to me than mine could have been to you, that I must and will write again. I saw your motive for not alluding to Natural Selection, and quite agreed in my mind in its wisdom. But at the same time it occurred to me that you might be giving it up, and that anyhow you could not safely allude to it without various "provisos" too long to give in a lecture. If I think continuously on some half-dozen structures of which we can at present see no use, I can persuade myself that Natural Selection is of quite subordinate importance. On the other hand, when I reflect on the innumerable structures, especially in plants, which twenty years ago would have been called simply "morphological" and useless, and which are now known to be highly important, I can persuade myself that every structure may have been developed through Natural Selection. It is really curious how many out of a list of structures which Bronn enumerated, as not possibly due to Natural Selection because of no functional importance, can now be shown to be highly important. Lobed leaves was, I believe, one case, and only two or three days ago Frank showed me how they act in a manner quite sufficiently important to account for the lobing of any large leaf. I am particularly delighted at what you say about domestic dogs, jackals, and wolves, because from mere indirect evidence I arrived in "Varieties of Domestic Animals" at exactly the same conclusion (299/3. Mr. Darwin's view was that domestic dogs descend from more than one wild species.) with respect to the domestic dogs of Europe and North America. See how important in another way this conclusion is; for no one can doubt that large and small dogs are perfectly fertile together, and produce fertile mongrels; and how well this supports the Pallasian doctrine (299/4. See Letter 80.) that domestication eliminates the sterility almost universal between forms slowly developed in a state of nature.

I humbly beg your pardon for bothering you with so long a note; but it is your own fault.

Plants are splendid for making one believe in Natural Selection, as will and consciousness are excluded. I have lately been experimenting on such a curious structure for bursting open the seed-coats: I declare one might as well say that a pair of scissors or nutcrackers had been developed through external conditions as the structure in question. (299/5. The peg or heel in Cucurbita: see "Power of Movement in Plants" page 102.)

LETTER 300. TO T.H. HUXLEY. Down, November 5th, 1880.

On reading over your excellent review (300/1. See "Nature," November 4th, 1880, page 1, a review of Volume I. of the publications of the "Challenger," to which Sir Wyville Thomson contributed a General Introduction.) with the sentence quoted from Sir Wyville Thomson, it seemed to me advisable, considering the nature of the publication, to notice "extreme variation" and another point. Now, will you read the enclosed, and if you approve, post it soon. If you disapprove, throw it in the fire, and thus add one more to the thousand kindnesses which you have done me. Do not write: I shall see result in next week's "Nature." Please observe that in the foul copy I had added a final sentence which I do not at first copy, as it seemed to me inferentially too contemptuous; but I have now pinned it to the back, and you can send it or not, as you think best,—that is, if you think any part worth sending. My request will not cost you much trouble—i.e. to read two pages, for I know that you can decide at once. I heartily enjoyed my talk with you on Sunday morning.

P.S.—If my manuscript appears too flat, too contemptuous, too spiteful, or too anything, I earnestly beseech you to throw it into the fire.

LETTER 301. CHARLES DARWIN TO THE EDITOR OF "NATURE."

(301/1. "Nature," November 11th, 1880, page 32.)

Down, November 5th, 1880.

Sir Wyville Thomson and Natural Selection.

I am sorry to find that Sir Wyville Thomson does not understand the principle of Natural Selection, as explained by Mr. Wallace and myself. If he had done so, he could not have written the following sentence in the Introduction to the Voyage of the "Challenger": "The character of the abyssal fauna refuses to give the least support to the theory which refers the evolution of species to extreme variation guided only by Natural Selection." This is a standard of criticism not uncommonly reached by theologians and metaphysicians, when they write on scientific subjects, but is something new as coming from a naturalist. Professor Huxley demurs to it in the last number of "Nature"; but he does not touch on the expression of extreme variation, nor on that of evolution being guided only by Natural Selection. Can Sir Wyville Thomson name any one who has said that the evolution of species depends only on Natural Selection? As far as concerns myself, I believe that no one has brought forward so many observations on the effects of the use and disuse of parts, as I have done in my "Variation of Animals and Plants under Domestication"; and these observations were made for this special object. I have likewise there adduced a considerable body of facts, showing the direct action of external conditions on organisms; though no doubt since my books were published much has been learnt on this head. If Sir Wyville Thomson were to visit the yard of a breeder, and saw all his cattle or sheep almost absolutely true—that is, closely similar, he would exclaim: "Sir, I see here no extreme variation; nor can I find any support to the belief that you have followed the principle of selection in the breeding of your animals." From what I formerly saw of breeders, I have no doubt that the man thus rebuked would have smiled and said not a word. If he had afterwards told the story to other breeders, I greatly fear that they would have used emphatic but irreverent language about naturalists.

(301/2. The following is the passage omitted by the advice of Huxley: see his "Life and Letters," II., page 14:—

"Perhaps it would have been wiser on my part to have remained quite silent, like the breeder; for, as Prof. Sedgwick remarked many years ago, in reference to the poor old Dean of York, who was never weary of inveighing against geologists, a man who talks about what he does not in the least understand, is invulnerable.")

LETTER 302. TO G.J. ROMANES.

(302/1. Part of this letter has been published in Mr. C. Barber's note on "Graft-Hybrids of the Sugar-Cane," in "The Sugar-Cane," November 1896.)

Down, January 1st, 1881.

I send the MS., but as far as I can judge by just skimming it, it will be of no use to you. It seems to bear on transitional forms. I feel sure that I have other and better cases, but I cannot remember where to look.

I should have written to you in a few days on the following case. The Baron de Villa Franca wrote to me from Brazil about two years ago, describing new varieties of sugar-cane which he had raised by planting two old varieties in apposition. I believe (but my memory is very faulty) that I wrote that I could not believe in such a result, and attributed the new varieties to the soil, etc. I believe that I did not understand what he meant by apposition. Yesterday a packet of MS. arrived from the Brazilian Legation, with a letter in French from Dr. Glass, Director of the Botanic Gardens, describing fully how he first attempted grafting varieties of sugar-cane in various ways, and always failed, and then split stems of two varieties, bound them together and planted them, and then raised some new and very valuable varieties, which, like crossed plants, seem to grow with extra vigour, are constant, and apparently partake of the character of the two varieties. The Baron also sends me an attested copy from a number of Brazilian cultivators of the success of the plan of raising new varieties. I am not sure whether the Brazilian Legation wishes me to return the document, but if I do not hear in three or four days that they must be returned, they shall be sent to you, for they seem to me well deserving your consideration.

Perhaps if I had been contented with my hyacinth bulbs being merely bound together without any true adhesion or rather growth together, I should have succeeded like the old Dutchman.

There is a deal of superfluous verbiage in the documents, but I have marked with pencil where the important part begins. The attestations are in duplicate. Now, after reading them will you give me your opinion whether the main parts are worthy of publication in "Nature": I am inclined to think so, and it is good to encourage science in out-of-the-way parts of the world.

Keep this note till you receive the documents or hear from me. I wonder whether two varieties of wheat could be similarly treated? No, I suppose not—from the want of lateral buds. I was extremely interested by your abstract on suicide.

LETTER 303. TO K. SEMPER. Down, February 6th, 1881.

Owing to all sorts of work, I have only just now finished reading your "Natural Conditions of Existence." (303/1. Semper's "Natural Conditions of Existence as they affect Animal Life" (International Science Series), 1881.) Although a book of small size, it contains an astonishing amount of matter, and I have been particularly struck with the originality with which you treat so many subjects, and at your scrupulous accuracy. In far the greater number of points I quite follow you in your conclusions, but I differ on some, and I suppose that no two men in the world would fully agree on so many different subjects. I have been interested on so many points, I can hardly say on which most. Perhaps as much on Geographical Distribution as on any other, especially in relation to M. Wagner. (No! no! about parasites interested me even more.) How strange that Wagner should have thought that I meant by struggle for existence, struggle for food. It is curious that he should not have thought of the endless adaptations for the dispersal of seeds and the fertilisation of flowers.

Again I was much interested about Branchipus and Artemia. (303/2. The reference is to Schmankewitsch's experiments, page 158: he kept Artemia salina in salt-water, gradually diluted with fresh-water until it became practically free from salt; the crustaceans gradually changed in the course of generations, until they acquired the characters of the genus Branchipus.) When I read imperfectly some years ago the original paper I could not avoid thinking that some special explanation would hereafter be found for so curious a case. I speculated whether a species very liable to repeated and great changes of conditions, might not acquire a fluctuating condition ready to be adapted to either conditions. With respect to Arctic animals being white (page 116 of your book) it might perhaps be worth your looking at what I say from Pallas' and my own observations in the "Descent of Man" (later editions) Chapter VIII., page 229, and Chapter XVIII., page 542.

I quite agree with what I gather to be your judgment, viz., that the direct action of the conditions of life on organisms, or the cause of their variability, is the most important of all subjects for the future. For some few years I have been thinking of commencing a set of experiments on plants, for they almost invariably vary when cultivated. I fancy that I see my way with the aid of continued self-fertilisation. But I am too old, and have not strength enough. Nevertheless the hope occasionally revives.

Finally let me thank you for the very kind manner in which you often refer to my works, and for the even still kinder manner in which you disagree with me.

With cordial thanks for the pleasure and instruction which I have derived from your book, etc.

LETTER 304. TO COUNT SAPORTA. Down, February 13th, 1881.

I received a week or two ago the work which you and Prof. Marion have been so kind as to send me. (304/1. Probably "L'Evolution du Regne vegetal," I. "Cryptogames," Saporta & Marion, Paris, 1881.) When it arrived I was much engaged, and this must be my excuse for not having sooner thanked you for it, and it will likewise account for my having as yet read only the preface.

But I now look forward with great pleasure to reading the whole immediately. If I then have any remarks worth sending, which is not very probable, I will write again. I am greatly pleased to see how boldly you express your belief in evolution, in the preface. I have sometimes thought that some of your countrymen have been a little timid in publishing their belief on this head, and have thus failed in aiding a good cause.

LETTER 305. TO R.G. WHITEMAN. Down, May 5th, 1881.

In the first edition of the "Origin," after the sentence ending with the words "...insects in the water," I added the following sentence:—

"Even in so extreme a case as this, if the supply of insects were constant, and if better adapted competitors did not already exist in the country, I can see no difficulty in a race of bears being rendered by Natural Selection more and more aquatic in their structures and habits, with larger and larger mouths, till a creature was produced as monstrous as a whale." (305/1. See Letters 110 and 120.)

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