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Coral Reefs
by Charles Darwin
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Captain Moresby informs me that he has seen a French chart of Diego Garcia made eighty years before his survey, and apparently very accurate; and from it he infers, that during this interval there has not been the smallest change in the depth on any of the knolls within the lagoon. It is also known that during the last fifty-one years, the eastern channel into the lagoon has neither become narrower, nor decreased in depth; and as there are numerous small knolls of living coral within it, some change might have been anticipated. Moreover, as the whole reef round the lagoon of this atoll has been converted into land—an unparalleled case, I believe, in an atoll of such large size,—and as the strip of land is for considerable spaces more than half a mile wide—also a very unusual circumstance,—we have the best possible evidence, that Diego Garcia has remained at its present level for a very long period. With this fact, and with the knowledge that no sensible change has taken place during eighty years in the coral-knolls, and considering that every single reef has reached the surface in other atolls, which do not present the smallest appearance of being older than Diego Garcia and Peros Banhos, and which are placed under the same external conditions with them, one is led to conclude that these submerged reefs, although covered with luxuriant coral, have no tendency to grow upwards, and that they would remain at their present levels for an almost indefinite period.

From the number of these knolls, from their position, size, and form, many of them being only one or two hundred yards across, with a rounded outline, and precipitous sides,—it is indisputable that they have been formed by the growth of coral; and this makes the case much more remarkable. In Peros Banhos and in the Great Chagos Bank, some of these almost columnar masses are 200 feet high, and their summits lie only from two to eight fathoms beneath the surface; therefore, a small proportional amount more of growth would cause them to attain the surface, like those numerous knolls, which rise from an equally great depth within the Maldiva atolls. We can hardly suppose that time has been wanting for the upward growth of the coral, whilst in Diego Garcia, the broad annular strip of land, formed by the continued accumulation of detritus, shows how long this atoll has remained at its present level. We must look to some other cause than the rate of growth; and I suspect it will be found in the reefs being formed of different species of corals, adapted to live at different depths.

The Great Chagos Bank is situated in the centre of the Chagos Group, and the Pitt and Speaker Banks at its two extreme points. These banks resemble atolls, except in their external rim being about eight fathoms submerged, and in being formed of dead rock, with very little living coral on it: a portion nine miles long of the annular reef of Peros Banhos atoll is in the same condition. These facts, as will hereafter be shown, render it very probable that the whole group at some former period subsided seven or eight fathoms; and that the coral perished on the outer margin of those atolls which are now submerged, but that it continued alive, and grew up to the surface on those which are now perfect. If these atolls did subside, and if from the suddenness of the movement or from any other cause, those corals which are better adapted to live at a certain depth than at the surface, once got possession of the knolls, supplanting the former occupants, they would exert little or no tendency to grow upwards. To illustrate this, I may observe, that if the corals of the upper zone on the outer edge of Keeling atoll were to perish, it is improbable that those of the lower zone would grow to the surface, and thus become exposed to conditions for which they do not appear to be adapted. The conjecture, that the corals on the submerged knolls within the Chagos atolls have analogous habits with those of the lower zone outside Keeling atoll, receives some support from a remark by Captain Moresby, namely, that they have a different appearance from those on the reefs in the Maldiva atolls, which, as we have seen, all rise to the surface: he compares the kind of difference to that of the vegetation under different climates. I have entered at considerable length into this case, although unable to throw much light on it, in order to show that an equal tendency to upward growth ought not to be attributed to all coral-reefs,—to those situated at different depths,—to those forming the ring of an atoll or those on the knolls within a lagoon,—to those in one area and those in another. The inference, therefore, that one reef could not grow up to the surface within a given time, because another, not known to be covered with the same species of corals, and not known to be placed under conditions exactly the same, has not within the same time reached the surface, is unsound.

SECTION 4.II.—ON THE RATE OF GROWTH OF CORAL-REEFS.

The remark made at the close of the last section, naturally leads to this division of our subject, which has not, I think, hitherto been considered under a right point of view. Ehrenberg (Ehrenberg, as before cited, pages 39, 46, and 50.) has stated, that in the Red Sea, the corals only coat other rocks in a layer from one to two feet in thickness, or at most to a fathom and a half; and he disbelieves that, in any case, they form, by their own proper growth, great masses, stratum over stratum. A nearly similar observation has been made by MM. Quoy and Gaimard ("Annales des Sciences Nat." tom. vi., page 28.), with respect to the thickness of some upraised beds of coral, which they examined at Timor and some other places. Ehrenberg (Ehrenberg, ut sup., page 42.) saw certain large massive corals in the Red Sea, which he imagines to be of such vast antiquity, that they might have been beheld by Pharaoh; and according to Mr. Lyell (Lyell's "Principles of Geology," book iii., chapter xviii.) there are certain corals at Bermuda, which are known by tradition, to have been living for centuries. To show how slowly coral-reefs grow upwards, Captain Beechey (Beechey's "Voyage to the Pacific," chapter viii.) has adduced the case of the Dolphin Reef off Tahiti, which has remained at the same depth beneath the surface, namely about two fathoms and a half, for a period of sixty-seven years. There are reefs in the Red Sea, which certainly do not appear (Ehrenberg, ut sup., page 43.) to have increased in dimensions during the last half-century, and from the comparison of old charts with recent surveys, probably not during the last two hundred years. These, and other similar facts, have so strongly impressed many with the belief of the extreme slowness of the growth of corals, that they have even doubted the possibility of islands in the great oceans having been formed by their agency. Others, again, who have not been overwhelmed by this difficulty, have admitted that it would require thousands, and tens of thousands of years, to form a mass, even of inconsiderable thickness; but the subject has not, I believe, been viewed in the proper light.

That masses of considerable thickness have been formed by the growth of coral, may be inferred with certainty from the following facts. In the deep lagoons of Peros Banhos and of the Great Chagos Bank, there are, as already described, small steep-sided knolls covered with living coral. There are similar knolls in the southern Maldiva atolls, some of which, as Captain Moresby assures me, are less than a hundred yards in diameter, and rise to the surface from a depth of between two hundred and fifty and three hundred feet. Considering their number, form, and position, it would be preposterous to suppose that they are based on pinnacles of any rock, not of coral formation; or that sediment could have been heaped up into such small and steep isolated cones. As no kind of living coral grows above the height of a few feet, we are compelled to suppose that these knolls have been formed by the successive growth and death of many individuals,—first one being broken off or killed by some accident, and then another, and one set of species being replaced by another set with different habits, as the reef rose nearer the surface, or as other changes supervened. The spaces between the corals would become filled up with fragments and sand, and such matter would probably soon be consolidated, for we learn from Lieutenant Nelson ("Geological Transactions," volume v., page 113.), that at Bermuda a process of this kind takes place beneath water, without the aid of evaporation. In reefs, also, of the barrier class, we may feel sure, as I have shown, that masses of great thickness have been formed by the growth of the coral; in the case of Vanikoro, judging only from the depth of the moat between the land and the reef, the wall of coral-rock must be at least three hundred feet in vertical thickness.

It is unfortunate that the upraised coral-islands in the Pacific have not been examined by a geologist. The cliffs of Elizabeth Island, in the Low Archipelago, are eighty feet high, and appear, from Captain Beechey's description, to consist of a homogeneous coral-rock. From the isolated position of this island, we may safely infer that it is an upraised atoll, and therefore that it has been formed by masses of coral, grown together. Savage Island seems, from the description of the younger Forster (Forster's "Voyage round the World with Cook," volume ii., pages 163, 167.), to have a similar structure, and its shores are about forty feet high: some of the Cook Islands also appear (Williams's "Narrative of Missionary Enterprise," page 30.) to be similarly composed. Captain Belcher, R.N., in a letter which Captain Beaufort showed me at the admiralty, speaking of Bow atoll, says, "I have succeeded in boring forty-five feet through coral-sand, when the auger became jammed by the falling in of the surrounding CREAMY matter." On one of the Maldiva atolls, Captain Moresby bored to a depth of twenty-six feet, when his auger also broke: he has had the kindness to give me the matter brought up; it is perfectly white, and like finely triturated coral-rock.

In my description of Keeling atoll, I have given some facts, which show that the reef probably has grown outwards; and I have found, just within the outer margin, the great mounds of Porites and of Millepora, with their summits lately killed, and their sides subsequently thickened by the growth of the coral: a layer, also, of Nullipora had already coated the dead surface. As the external slope of the reef is the same round the whole of this atoll, and round many other atolls, the angle of inclination must result from an adaption between the growing powers of the coral, and the force of the breakers, and their action on the loose sediment. The reef, therefore, could not increase outwards, without a nearly equal addition to every part of the slope, so that the original inclination might be preserved, and this would require a large amount of sediment, all derived from the wear of corals and shells, to be added to the lower part. Moreover, at Keeling atoll, and probably in many other cases, the different kinds of corals would have to encroach on each other; thus the Nulliporae cannot increase outwards without encroaching on the Porites and Millepora complanata, as is now taking place; nor these latter without encroaching on the strongly branched Madreporet, the Millepora alcicornis, and some Astraeas; nor these again without a foundation being formed for them within the requisite depth, by the accumulation of sediment. How slow, then, must be the ordinary lateral or outward growth of such reefs. But off Christmas atoll, where the sea is much more shallow than is usual, we have good reason to believe that, within a period not very remote, the reef has increased considerably in width. The land has the extraordinary breadth of three miles; it consists of parallel ridges of shells and broken corals, which furnish "an incontestable proof," as observed by Cook (Cook's "Third Voyage," book III., chapter x.), "that the island has been produced by accessions from the sea, and is in a state of increase." The land is fronted by a coral-reef, and from the manner in which islets are known to be formed, we may feel confident that the reef was not three miles wide, when the first, or most backward ridge, was thrown up; and, therefore, we must conclude that the reef has grown outwards during the accumulation of the successive ridges. Here then, a wall of coral-rock of very considerable breadth has been formed by the outward growth of the living margin, within a period during which ridges of shells and corals, lying on the bare surface, have not decayed. There can be little doubt, from the account given by Captain Beechey, that Matilda atoll, in the Low Archipelago, has been converted in the space of thirty-four years, from being, as described by the crew of a wrecked whaling vessel, a "reef of rocks" into a lagoon-island, fourteen miles in length, with "one of its sides covered nearly the whole way with high trees." (Beechey's "Voyage to the Pacific," chapter vii. and viii.) The islets, also, on Keeling atoll, it has been shown, have increased in length, and since the construction of an old chart, several of them have become united into one long islet; but in this case, and in that of Matilda atoll, we have no proof, and can only infer as probable, that the reef, that is the foundation of the islets, has increased as well as the islets themselves.

After these considerations, I attach little importance, as indicating the ordinary and still less the possible rate of OUTWARD growth of coral-reefs, to the fact that certain reefs in the Red Sea have not increased during a long interval of time; or to other such cases, as that of Ouluthy atoll in the Caroline group, where every islet, described a thousand years before by Cantova was found in the same state by Lutke (F. Lutke's "Voyage autour du Monde." In the group Elato, however, it appears that what is now the islet Falipi, is called in Cantova's Chart, the Banc de Falipi. It is not stated whether this has been caused by the growth of coral, or by the accumulation of sand.),—without it could be shown that, in these cases, the conditions were favourable to the vigorous and unopposed growth of the corals living in the different zones of depth, and that a proper basis for the extent of the reef was present. The former conditions must depend on many contingencies, and in the deep oceans where coral formations most abound, a basis within the requisite depth can rarely be present.

Nor do I attach any importance to the fact of certain submerged reefs, as those off Tahiti, or those within Diego Garcia not now being nearer the surface than they were many years ago, as an indication of the rate under favourable circumstances of the UPWARD growth of reefs; after it has been shown, that all the reefs have grown to the surface in some of the Chagos atolls, but that in neighbouring atolls which appear to be of equal antiquity and to be exposed to the same external conditions, every reef remains submerged; for we are almost driven to attribute this to a difference, not in the rate of growth, but in the habits of the corals in the two cases.

In an old-standing reef, the corals, which are so different in kind on different parts of it, are probably all adapted to the stations they occupy, and hold their places, like other organic beings, by a struggle one with another, and with external nature; hence we may infer that their growth would generally be slow, except under peculiarly favourable circumstances. Almost the only natural condition, allowing a quick upward growth of the whole surface of a reef, would be a slow subsidence of the area in which it stood; if, for instance, Keeling atoll were to subside two or three feet, can we doubt that the projecting margin of live coral, about half an inch in thickness, which surrounds the dead upper surfaces of the mounds of Porites, would in this case form a concentric layer over them, and the reef thus increase upwards, instead of, as at present, outwards? The Nulliporae are now encroaching on the Porites and Millepora, but in this case might we not confidently expect that the latter would, in their turn, encroach on the Nulliporae? After a subsidence of this kind, the sea would gain on the islets, and the great fields of dead but upright corals in the lagoon, would be covered by a sheet of clear water; and might we not then expect that these reefs would rise to the surface, as they anciently did when the lagoon was less confined by islets, and as they did within a period of ten years in the schooner-channel, cut by the inhabitants? In one of the Maldiva atolls, a reef, which within a very few years existed as an islet bearing cocoa-nut trees, was found by Lieutenant Prentice "ENTIRELY COVERED WITH LIVE CORAL AND MADREPORE." The natives believe that the islet was washed away by a change in the currents, but if, instead of this, it had quietly subsided, surely every part of the island which offered a solid foundation, would in a like manner have become coated with living coral.

Through steps such as these, any thickness of rock, composed of a singular intermixture of various kinds of corals, shells, and calcareous sediment, might be formed; but without subsidence, the thickness would necessarily be determined by the depth at which the reef-building polypifers can exist. If it be asked, at what rate in years I suppose a reef of coral favourably circumstanced could grow up from a given depth; I should answer, that we have no precise evidence on this point, and comparatively little concern with it. We see, in innumerable points over wide areas, that the rate has been sufficient, either to bring up the reefs from various depths to the surface, or, as is more probable, to keep them at the surface, during progressive subsidences; and this is a much more important standard of comparison than any cycle of years.

It may, however, be inferred from the following facts, that the rate in years under favourable circumstances would be very far from slow. Dr. Allan, of Forres, has, in his MS. Thesis deposited in the library of the Edinburgh University (extracts from which I owe to the kindness of Dr. Malcolmson), the following account of some experiments, which he tried during his travels in the years 1830 to 1832 on the east coast of Madagascar. "To ascertain the rise and progress of the coral-family, and fix the number of species met with at Foul Point (latitude 17 deg 40') twenty species of coral were taken off the reef and planted apart on a sand-bank THREE FEET DEEP AT LOW WATER. Each portion weighed ten pounds, and was kept in its place by stakes. Similar quantities were placed in a clump and secured as the rest. This was done in December 1830. In July following, each detached mass was nearly level with the sea at low water, quite immovable, and several feet long, stretching as the parent reef, with the coast current from north to south. The masses accumulated in a clump were found equally increased, but some of the species in such unequal ratios, as to be growing over each other." The loss of Dr. Allan's magnificent collection by shipwreck, unfortunately prevents its being known to what genera these corals belonged; but from the numbers experimented on, it is certain that all the more conspicuous kinds must have been included. Dr. Allan informs me, in a letter, that he believes it was a Madrepora, which grew most vigorously. One may be permitted to suspect that the level of the sea might possibly have been somewhat different at the two stated periods; nevertheless, it is quite evident that the growth of the ten-pound masses, during the six or seven months, at the end of which they were found immovably fixed (It is stated by De la Beche ("Geological Manual," page 143), on the authority of Mr. Lloyd, who surveyed the Isthmus of Panama, that some specimens of Polypifers, placed by him in a sheltered pool of water, were found in the course of a few days firmly fixed by the secretion of a stony matter, to the bottom) and several feet in length, must have been very great. The fact of the different kinds of coral, when placed in one clump, having increased in extremely unequal ratios, is very interesting, as it shows the manner in which a reef, supporting many species of coral, would probably be affected by a change in the external conditions favouring one kind more than another. The growth of the masses of coral in N. and S. lines parallel to the prevailing currents, whether due to the drifting of sediment or to the simple movement of the water, is, also, a very interesting circumstance.

A fact, communicated to me by Lieutenant Wellstead, I.N., in some degree corroborates the result of Dr. Allan's experiments: it is, that in the Persian Gulf a ship had her copper bottom encrusted in the course of twenty months with a layer of coral, TWO FEET in thickness, which it required great force to remove, when the vessel was docked: it was not ascertained to what order this coral belonged. The case of the schooner-channel choked up with coral in an interval of less than ten years, in the lagoon of Keeling atoll, should be here borne in mind. We may also infer, from the trouble which the inhabitants of the Maldiva atolls take to root out, as they express it, the coral-knolls from their harbours, that their growth can hardly be very slow. (Mr. Stutchbury ("West of England Journal", No. I., page 50.) has described a specimen of Agaricia, "weighing 2 lbs. 9 oz., which surrounds a species of oyster, whose age could not be more than two years, and yet is completely enveloped by this dense coral." I presume that the oyster was living when the specimen was procured; otherwise the fact tells nothing. Mr. Stutchbury also mentions an anchor, which had become entirely encrusted with coral in fifty years; other cases, however, are recorded of anchors which have long remained amidst coral-reefs without having become coated. The anchor of the "Beagle", in 1832, after having been down exactly one month at Rio de Janeiro, was so thickly coated by two species of Tubularia, that large spaces of the iron were entirely concealed; the tufts of this horny zoophyte were between two and three inches in length. It has been attempted to compute, but I believe erroneously, the rate of growth of a reef, from the fact mentioned by Captain Beechey, of the Chama gigas being embedded in coral-rock. But it should be remembered, that some species of this genus invariably live, both whilst young and old, in cavities, which the animal has the power of enlarging with its growth. I saw many of these shells thus embedded in the outer "flat" of Keeling atoll, which is composed of dead rock; and therefore the cavities in this case had no relation whatever with the growth of coral. M. Lesson, also, speaking of this shell (Partie Zoolog. "Voyage de la 'Coquille'"), has remarked, "que constamment ses valves etaient engages completement dans la masse des Madrepores.")

From the facts given in this section, it may be concluded, first, that considerable thicknesses of rock have certainly been formed within the present geological area by the growth of coral and the accumulation of its detritus; and, secondly, that the increase of individual corals and of reefs, both outwards or horizontally and upwards or vertically, under the peculiar conditions favourable to such increase, is not slow, when referred either to the standard of the average oscillations of level in the earth's crust, or to the more precise but less important one of a cycle of years.

SECTION 4.III.—ON THE DEPTHS AT WHICH REEF-BUILDING POLYPIFERS CAN LIVE.

I have already described in detail, which might have appeared trivial, the nature of the bottom of the sea immediately surrounding Keeling atoll; and I will now describe with almost equal care the soundings off the fringing-reefs of Mauritius. I have preferred this arrangement, for the sake of grouping together facts of a similar nature. I sounded with the wide bell-shaped lead which Captain Fitzroy used at Keeling Island, but my examination of the bottom was confined to a few miles of coast (between Port Louis and Tomb Bay) on the leeward side of the island. The edge of the reef is formed of great shapeless masses of branching Madrepores, which chiefly consist of two species,—apparently M. corymbosa and pocillifera,— mingled with a few other kinds of coral. These masses are separated from each other by the most irregular gullies and cavities, into which the lead sinks many feet. Outside this irregular border of Madrepores, the water deepens gradually to twenty fathoms, which depth generally is found at the distance of from half to three-quarters of a mile from the reef. A little further out the depth is thirty fathoms, and thence the bank slopes rapidly into the depths of the ocean. This inclination is very gentle compared with that outside Keeling and other atolls, but compared with most coasts it is steep. The water was so clear outside the reef, that I could distinguish every object forming the rugged bottom. In this part, and to a depth of eight fathoms, I sounded repeatedly, and at each cast pounded the bottom with the broad lead, nevertheless the arming invariably came up perfectly clean, but deeply indented. From eight to fifteen fathoms a little calcareous sand was occasionally brought up, but more frequently the arming was simply indented. In all this space the two Madrepores above mentioned, and two species of Astraea, with rather large stars, seemed the commonest kinds (Since the preceding pages were printed off, I have received from Mr. Lyell a very interesting pamphlet, entitled "Remarks upon Coral Formations," etc., by J. Couthouy, Boston, United States, 1842. There is a statement (page 6), on the authority of the Rev. J. Williams, corroborating the remarks made by Ehrenberg and Lyell (page 71 of this volume), on the antiquity of certain individual corals in the Red Sea and at Bermuda; namely, that at Upolu, one of the Navigator Islands, "particular clumps of coral are known to the fishermen by name, derived from either some particular configuration or tradition attached to them, and handed down from time immemorial." With respect to the thickness of masses of coral-rock, it clearly appears, from the descriptions given by Mr. Couthouy (pages 34, 58) that Mangaia and Aurora Islands are upraised atolls, composed of coral rock: the level summit of the former is about three hundred feet, and that of Aurora Island is two hundred feet above the sea-level.); and it must be noticed that twice at the depth of fifteen fathoms, the arming was marked with a clean impression of an Astraea. Besides these lithophytes, some fragments of the Millepora alcicornis, which occurs in the same relative position at Keeling Island, were brought up; and in the deeper parts there were large beds of a Seriatopora, different from S. subulata, but closely allied to it. On the beach within the reef, the rolled fragments consisted chiefly of the corals just mentioned, and of a massive Porites, like that at Keeling atoll, of a Meandrina, Pocillopora verrucosa, and of numerous fragments of Nullipora. From fifteen to twenty fathoms the bottom was, with few exceptions, either formed of sand, or thickly covered with Seriatopora: this delicate coral seems to form at these depths extensive beds unmingled with any other kind. At twenty fathoms, one sounding brought up a fragment of Madrepora apparently M. pocillifera, and I believe it is the same species (for I neglected to bring specimens from both stations) which mainly forms the upper margin of the reef; if so, it grows in depths varying from 0 to 20 fathoms. Between 20 and 23 fathoms I obtained several soundings, and they all showed a sandy bottom, with one exception at 30 fathoms, when the arming came up scooped out, as if by the margin of a large Caryophyllia. Beyond 33 fathoms I sounded only once; and from 86 fathoms, at the distance of one mile and a third from the edge of the reef, the arming brought up calcareous sand with a pebble of volcanic rock. The circumstance of the arming having invariably come up quite clean, when sounding within a certain number of fathoms off the reefs of Mauritius and Keeling atoll (eight fathoms in the former case, and twelve in the latter) and of its having always come up (with one exception) smoothed and covered with sand, when the depth exceeded twenty fathoms, probably indicates a criterion, by which the limits of the vigorous growth of coral might in all cases be readily ascertained. I do not, however, suppose that if a vast number of soundings were obtained round these islands, the limit above assigned would be found never to vary, but I conceive the facts are sufficient to show, that the exceptions would be few. The circumstance of a GRADUAL change, in the two cases, from a field of clean coral to a smooth sandy bottom, is far more important in indicating the depth at which the larger kinds of coral flourish than almost any number of separate observations on the depth, at which certain species have been dredged up. For we can understand the gradation, only as a prolonged struggle against unfavourable conditions. If a person were to find the soil clothed with turf on the banks of a stream of water, but on going to some distance on one side of it, he observed the blades of grass growing thinner and thinner, with intervening patches of sand, until he entered a desert of sand, he would safely conclude, especially if changes of the same kind were noticed in other places, that the presence of the water was absolutely necessary to the formation of a thick bed of turf: so may we conclude, with the same feeling of certainty, that thick beds of coral are formed only at small depths beneath the surface of the sea.

I have endeavoured to collect every fact, which might either invalidate or corroborate this conclusion. Captain Moresby, whose opportunities for observation during his survey of the Maldiva and Chagos Archipelagoes have been unrivalled, informs me, that the upper part or zone of the steep-sided reefs, on the inner and outer coasts of the atolls in both groups, invariably consists of coral, and the lower parts of sand. At seven or eight fathoms depth, the bottom is formed, as could be seen through the clear water, of great living masses of coral, which at about ten fathoms generally stand some way apart from each other, with patches of white sand between them, and at a little greater depth these patches become united into a smooth steep slope, without any coral. Captain Moresby, also, informs me in support of his statement, that he found only decayed coral on the Padua Bank (northern part of the Laccadive group) which has an average depth between twenty-five and thirty-five fathoms, but that on some other banks in the same group with only ten or twelve fathoms water on them (for instance, the Tillacapeni bank), the coral was living.

With regard to the coral-reefs in the Red Sea, Ehrenberg has the following passage:—"The living corals do not descend there into great depths. On the edges of islets and near reefs, where the depth was small, very many lived; but we found no more even at six fathoms. The pearl-fishers at Yemen and Massaua asserted that there was no coral near the pearl-banks at nine fathoms depth, but only sand. We were not able to institute any more special researches." (Ehrenberg, "Uber die Natur," etc., page 50.) I am, however, assured both by Captain Moresby and Lieutenant Wellstead, that in the more northern parts of the Red Sea, there are extensive beds of living coral at a depth of twenty-five fathoms, in which the anchors of their vessels were frequently entangled. Captain Moresby attributes the less depth, at which the corals are able to live in the places mentioned by Ehrenberg, to the greater quantity of sediment there; and the situations, where they were flourishing at the depth of twenty-five fathoms, were protected, and the water was extraordinarily limpid. On the leeward side of Mauritius where I found the coral growing at a somewhat greater depth than at Keeling atoll, the sea, owing apparently to its tranquil state, was likewise very clear. Within the lagoons of some of the Marshall atolls, where the water can be but little agitated, there are, according to Kotzebue, living beds of coral in twenty-five fathoms. From these facts, and considering the manner in which the beds of clean coral off Mauritius, Keeling Island, the Maldiva and Chagos atolls, graduated into a sandy slope, it appears very probable that the depth, at which reef-building polypifers can exist, is partly determined by the extent of inclined surface, which the currents of the sea and the recoiling waves have the power to keep free from sediment.

MM. Quoy and Gaimard ("Annales des Sci. Nat." tom. vi.) believe that the growth of coral is confined within very limited depths; and they state that they never found any fragment of an Astraea (the genus they consider most efficient in forming reefs) at a depth above twenty-five or thirty feet. But we have seen that in several places the bottom of the sea is paved with massive corals at more than twice this depth; and at fifteen fathoms (or twice this depth) off the reefs of Mauritius, the arming was marked with the distinct impression of a living Astraea. Millepora alcicornis lives in from 0 to 12 fathoms, and the genera Madrepora and Seriatopora from 0 to 20 fathoms. Captain Moresby has given me a specimen of Sideropora scabra (Porites of Lamarck) brought up alive from 17 fathoms. Mr. Couthouy ("Remarks on Coral Formations," page 12.) states that he has dredged up on the Bahama banks considerable masses of Meandrina from 16 fathoms, and he has seen this coral growing in 20 fathoms. A Caryophyllia, half an inch in diameter, was dredged up alive from 80 fathoms off Juan Fernandez (latitude 33 deg S.) by Captain P.P. King (I am indebted to Mr. Stokes for having kindly communicated this fact to me, together with much other valuable information.): this is the most remarkable fact with which I am acquainted, showing the depth at which a genus of corals often found on reefs, can exist.

We ought, however, to feel less surprise at this fact, as Caryophyllia alone of the lamelliform genera, ranges far beyond the tropics; it is found in Zetland (Fleming's "British Animals," genus Caryophyllia.) in Latitude 60 deg N. in deep water, and I procured a small species from Tierra del Fuego in Latitude 53 deg S. Captain Beechey informs me, that branches of pink and yellow coral were frequently brought up from between twenty and twenty-five fathoms off the Low atolls; and Lieutenant Stokes, writing to me from the N.W. coast of Australia, says that a strongly branched coral was procured there from thirty fathoms; unfortunately it is not known to what genera these corals belong.

(I will record in the form of a note all the facts that I have been able to collect on the depths, both within and without the tropics, at which those corals and corallines can live, which there is no reason to suppose ever materially aid in the construction of a reef.

(In the following list the name of the Zoophyte is followed by the depth in fathoms, the country and degrees S. latitude, and the authority. Where no authority is given, the observation is Darwin's own.)

SERTULARIA, 40, Cape Horn 66.

CELLARIA, 40, Cape Horn 66.

CELLARIA, A minute scarlet encrusting species, found living, 190, Keeling Atoll, 12.

CELLARIA, An allied, small stony sub-generic form, 48, St Cruz Riv. 50.

A coral allied to VINCULARIA, with eight rows of cells, 40, Cape Horn.

TUBULIPORA, near to T. patima, 40, Cape Horn.

TUBULIPORA, near to T. patima, 94, East Chiloe 43.

CELLEPORA, several species, and allied sub-generic forms, 40, Cape Horn.

CELLEPORA, several species, and allied sub-generic forms, 40 and 57, Chonos Archipelago 45.

CELLEPORA, several species, and allied sub-generic forms, 48, St Cruz 50.

ESCHARA, 30, Tierra del Fuego 53.

ESCHARA, 48, St Cruz R. 50.

RETEPORA, 40, Cape Horn.

RETEPORA, 100, Cape of Good Hope 34, Quoy and Gaimard, "Ann. Scien. Nat." tome vi., page 284.

MILLEPORA, a strong coral with cylindrical branches, of a pink colour, about two inches high, resembling in the form of its orifices M. aspera of Lamarck, 94 and 30, E. Chiloe 43, Tierra del Fuego 53.

CORALIUM, 120, Barbary 33 N., Peyssonel in paper read to Royal Society May 1752.

ANTIPATHES, 16, Chonos 45.

GORGONIA (or an allied form), 160, Abrolhos on the coast of Brazil 18, Captain Beechey informed me of this fact in a letter.

Ellis ("Nat. Hist. of Coralline," page 96) states that Ombellularia was procured in latitude 79 deg N. STICKING to a LINE from the depth of 236 fathoms; hence this coral either must have been floating loose, or was entangled in stray line at the bottom. Off Keeling atoll a compound Ascidia (Sigillina) was brought up from 39 fathoms, and a piece of sponge, apparently living, from 70, and a fragment of Nullipora also apparently living from 92 fathoms. At a greater depth than 90 fathoms off this coral island, the bottom was thickly strewed with joints of Halimeda and small fragments of other Nulliporae, but all dead. Captain B. Allen, R.N., informs me that in the survey of the West Indies it was noticed that between the depth of 10 and 200 fathoms, the sounding lead very generally came up coated with the dead joints of a Halimeda, of which he showed me specimens. Off Pernambuco, in Brazil, in about twelve fathoms, the bottom was covered with fragments dead and alive of a dull red Nullipora, and I infer from Roussin's chart, that a bottom of this kind extends over a wide area. On the beach, within the coral-reefs of Mauritius, vast quantities of fragments of Nulliporae were piled up. From these facts it appears, that these simply organized bodies are amongst the most abundant productions of the sea.)

Although the limit of depth, at which each particular kind of coral ceases to exist, is far from being accurately known; yet when we bear in mind the manner in which the clumps of coral gradually became infrequent at about the same depth, and wholly disappeared at a greater depth than twenty fathoms, on the slope round Keeling atoll, on the leeward side of the Mauritius, and at rather less depth, both without and within the atolls of the Maldiva and Chagos Archipelagoes; and when we know that the reefs round these islands do not differ from other coral formations in their form and structure, we may, I think, conclude that in ordinary cases, reef-building polypifers do not flourish at greater depths than between twenty and thirty fathoms.

It has been argued ("Journal of the Royal Geographical Society," 1831, page 218.) that reefs may possibly rise from very great depths through the means of small corals, first making a platform for the growth of the stronger kinds. This, however, is an arbitrary supposition: it is not always remembered, that in such cases there is an antagonist power in action, namely, the decay of organic bodies, when not protected by a covering of sediment, or by their own rapid growth. We have, moreover, no right to calculate on unlimited time for the accumulation of small organic bodies into great masses. Every fact in geology proclaims that neither the land, nor the bed of the sea retain for indefinite periods the same level. As well might it be imagined that the British Seas would in time become choked up with beds of oysters, or that the numerous small corallines off the inhospitable shores of Tierra del Fuego would in time form a solid and extensive coral-reef.

CHAPTER V.—THEORY OF THE FORMATION OF THE DIFFERENT CLASSES OF CORAL-REEFS.

The atolls of the larger archipelagoes are not formed on submerged craters, or on banks of sediment.—Immense areas interspersed with atolls.—Their subsidence.—The effects of storms and earthquakes on atolls.—Recent changes in their state.—The origin of barrier-reefs and of atolls.—Their relative forms.—The step-formed ledges and walls round the shores of some lagoons.—The ring-formed reefs of the Maldiva atolls.—The submerged condition of parts or of the whole of some annular reefs.—The disseverment of large atolls.—The union of atolls by linear reefs.—The Great Chagos Bank.—Objections from the area and amount of subsidence required by the theory, considered.—The probable composition of the lower parts of atolls.

The naturalists who have visited the Pacific, seem to have had their attention riveted by the lagoon-islands, or atolls,—those singular rings of coral-land which rise abruptly out of the unfathomable ocean—and have passed over, almost unnoticed, the scarcely less wonderful encircling barrier-reefs. The theory most generally received on the formation of atolls, is that they are based on submarine craters; but where can we find a crater of the shape of Bow atoll, which is five times as long as it is broad (Plate I., Figure 4); or like that of Menchikoff Island (Plate II., Figure 3.), with its three loops, together sixty miles in length; or like Rimsky Korsacoff, narrow, crooked, and fifty-four miles long; or like the northern Maldiva atolls, made up of numerous ring-formed reefs, placed on the margin of a disc,—one of which discs is eighty-eight miles in length, and only from ten to twenty in breadth? It is, also, not a little improbable, that there should have existed as many craters of immense size crowded together beneath the sea, as there are now in some parts atolls. But this theory lies under a greater difficulty, as will be evident, when we consider on what foundations the atolls of the larger archipelagoes rest: nevertheless, if the rim of a crater afforded a basis at the proper depth, I am far from denying that a reef like a perfectly characterised atoll might not be formed; some such, perhaps, now exist; but I cannot believe in the possibility of the greater number having thus originated.

An earlier and better theory was proposed by Chamisso (Kotzebue's "First Voyage," volume iii., page 331.); he supposes that as the more massive kinds of corals prefer the surf, the outer portions, in a reef rising from a submarine basis, would first reach the surface and consequently form a ring. But on this view it must be assumed, that in every case the basis consists of a flat bank; for if it were conically formed, like a mountainous mass, we can see no reason why the coral should spring up from the flanks, instead of from the central and highest parts: considering the number of the atolls in the Pacific and Indian Oceans, this assumption is very improbable. As the lagoons of atolls are sometimes even more than forty fathoms deep, it must, also, be assumed on this view, that at a depth at which the waves do not break, the coral grows more vigorously on the edges of a bank than on its central part; and this is an assumption without any evidence in support of it. I remarked, in the third chapter, that a reef, growing on a detached bank, would tend to assume an atoll-like structure; if, therefore, corals were to grow up from a bank, with a level surface some fathoms submerged, having steep sides and being situated in a deep sea, a reef not to be distinguished from an atoll, might be formed: I believe some such exist in the West Indies. But a difficulty of the same kind with that affecting the crater theory, runners, as we shall presently see, this view inapplicable to the greater number of atolls.

No theory worthy of notice has been advanced to account for those barrier-reefs, which encircle islands of moderate dimensions. The great reef which fronts the coast of Australia has been supposed, but without any special facts, to rest on the edge of a submarine precipice, extending parallel to the shore. The origin of the third class or of fringing-reefs presents, I believe, scarcely any difficulty, and is simply consequent on the polypifers not growing up from great depths, and their not flourishing close to gently shelving beaches where the water is often turbid.

What cause, then, has given to atolls and barrier-reefs their characteristic forms? Let us see whether an important deduction will not follow from the consideration of these two circumstances, first, the reef-building corals flourishing only at limited depths; and secondly, the vastness of the areas interspersed with coral-reefs and coral-islets, none of which rise to a greater height above the level of the sea, than that attained by matter thrown up by the waves and winds. I do not make this latter statement vaguely; I have carefully sought for descriptions of every island in the intertropical seas; and my task has been in some degree abridged by a map of the Pacific, corrected in 1834 by MM. D'Urville and Lottin, in which the low islands are distinguished from the high ones (even from those much less than a hundred feet in height) by being written without a capital letter; I have detected a few errors in this map, respecting the height of some of the islands, which will be noticed in the Appendix, where I treat of coral formations in geographical order. To the Appendix, also, I must refer for a more particular account of the data on which the statements on the next page are grounded. I have ascertained, and chiefly from the writings of Cook, Kotzebue, Bellinghausen, Duperrey, Beechey, and Lutke, regarding the Pacific; and from Moresby (See also Captain Owen's and Lieutenant Wood's papers in the "Geographical Journal", on the Maldiva and Laccadive Archipelagoes. These officers particularly refer to the lowness of the islets; but I chiefly ground my assertion respecting these two groups, and the Chagos group, from information communicated to me by Captain Moresby.) with respect to the Indian Ocean, that in the following cases the term "low island" strictly means land of the height commonly attained by matter thrown up by the winds and the waves of an open sea. If we draw a line (the plan I have always adopted) joining the external atolls of that part of the Low Archipelago in which the islands are numerous, the figure will be a pointed ellipse (reaching from Hood to Lazaref Island), of which the longer axis is 840 geographical miles, and the shorter 420 miles; in this space (I find from Mr. Couthouy's pamphlet (page 58) that Aurora Island is about two hundred feet in height; it consists of coral-rock, and seems to have been formed by the elevation of an atoll. It lies north-east of Tahiti, close without the line bounding the space coloured dark blue in the map appended to this volume. Honden Island, which is situated in the extreme north-west part of the Low Archipelago, according to measurements made on board the "Beagle", whilst sailing by, is 114 feet from the SUMMIT OF THE TREES to the water's edge. This island appeared to resemble the other atolls of the group.) none of the innumerable islets united into great rings rise above the stated level. The Gilbert group is very narrow, and 300 miles in length. In a prolonged line from this group, at the distance of 240 miles, is the Marshall Archipelago, the figure of which is an irregular square, one end being broader than the other; its length is 520 miles, with an average width of 240; these two groups together are 1,040 miles in length, and all their islets are low. Between the southern end of the Gilbert and the northern end of Low Archipelago, the ocean is thinly strewed with islands, all of which, as far as I have been able to ascertain, are low; so that from nearly the southern end of the Low Archipelago, to the northern end of the Marshall Archipelago, there is a narrow band of ocean, more than 4,000 miles in length, containing a great number of islands, all of which are low. In the western part of the Caroline Archipelago, there is a space of 480 miles in length, and about 100 broad, thinly interspersed with low islands. Lastly, in the Indian Ocean, the archipelago of the Maldivas is 470 miles in length, and 60 in breadth; that of the Laccadives is 150 by 100 miles; as there is a low island between these two groups, they may be considered as one group of 1,000 miles in length. To this may be added the Chagos group of low islands, situated 280 miles distant, in a line prolonged from the southern extremity of the Maldivas. This group, including the submerged banks, is 170 miles in length and 80 in breadth. So striking is the uniformity in direction of these three archipelagoes, all the islands of which are low, that Captain Moresby, in one of his papers, speaks of them as parts of one great chain, nearly 1,500 miles long. I am, then, fully justified in repeating, that enormous spaces, both in the Pacific and Indian Oceans, are interspersed with islands, of which not one rises above that height, to which the waves and winds in an open sea can heap up matter.

On what foundations, then, have these reefs and islets of coral been constructed? A foundation must originally have been present beneath each atoll at that limited depth, which is indispensable for the first growth of the reef-building polypifers. A conjecture will perhaps be hazarded, that the requisite bases might have been afforded by the accumulation of great banks of sediment, which owing to the action of superficial currents (aided possibly by the undulatory movement of the sea) did not quite reach the surface,—as actually appears to have been the case in some parts of the West Indian Sea. But in the form and disposition of the groups of atolls, there is nothing to countenance this notion; and the assumption without any proof, that a number of immense piles of sediment have been heaped on the floor of the great Pacific and Indian Oceans, in their central parts far remote from land, and where the dark blue colour of the limpid water bespeaks its purity, cannot for one moment be admitted.

The many widely-scattered atolls must, therefore, rest on rocky bases. But we cannot believe that the broad summit of a mountain lies buried at the depth of a few fathoms beneath every atoll, and nevertheless throughout the immense areas above-named, with not one point of rock projecting above the level of the sea; for we may judge with some accuracy of mountains beneath the sea, by those on the land; and where can we find a single chain several hundred miles in length and of considerable breadth, much less several such chains, with their many broad summits attaining the same height, within from 120 to 180 feet? If the data be thought insufficient, on which I have grounded my belief, respecting the depth at which the reef-building polypifers can exist, and it be assumed that they can flourish at a depth of even one hundred fathoms, yet the weight of the above argument is but little diminished, for it is almost equally improbable, that as many submarine mountains, as there are low islands in the several great and widely separated areas above specified, should all rise within six hundred feet of the surface of the sea and not one above it, as that they should be of the same height within the smaller limit of one or two hundred feet. So highly improbable is this supposition, that we are compelled to believe, that the bases of the many atolls did never at any one period all lie submerged within the depth of a few fathoms beneath the surface, but that they were brought into the requisite position or level, some at one period and some at another, through movements in the earth's crust. But this could not have been effected by elevation, for the belief that points so numerous and so widely separated were successively uplifted to a certain level, but that not one point was raised above that level, is quite as improbable as the former supposition, and indeed differs little from it. It will probably occur to those who have read Ehrenberg's account of the Reefs of the Red Sea, that many points in these great areas may have been elevated, but that as soon as raised, the protuberant parts were cut off by the destroying action of the waves: a moment's reflection, however, on the basin-like form of the atolls, will show that this is impossible; for the upheaval and subsequent abrasion of an island would leave a flat disc, which might become coated with coral, but not a deeply concave surface; moreover, we should expect to see, in some parts at least, the rock of the foundation brought to the surface. If, then, the foundations of the many atolls were not uplifted into the requisite position, they must of necessity have subsided into it; and this at once solves every difficulty (The additional difficulty on the crater hypothesis before alluded to, will now be evident; for on this view the volcanic action must be supposed to have formed within the areas specified a vast number of craters, all rising within a few fathoms of the surface, and not one above it. The supposition that the craters were at different times upraised above the surface, and were there abraded by the surf and subsequently coated by corals, is subject to nearly the same objections with those given above in this paragraph; but I consider it superfluous to detail all the arguments opposed to such a notion. Chamisso's theory, from assuming the existence of so many banks, all lying at the proper depth beneath the water, is also vitally defective. The same observation applies to an hypothesis of Lieutenant Nelson's ("Geolog. Trans." volume v., page 122), who supposes that the ring-formed structure is caused by a greater number of germs of corals becoming attached to the declivity, than to the central plateau of a submarine bank: it likewise applies to the notion formerly entertained (Forster's "Observ." page 151), that lagoon-islands owe their peculiar form to the instinctive tendencies of the polypifers. According to this latter view, the corals on the outer margin of the reef instinctively expose themselves to the surf in order to afford protection to corals living in the lagoon, which belong to other genera, and to other families!), for we may safely infer, from the facts given in the last chapter, that during a gradual subsidence the corals would be favourably circumstanced for building up their solid frame works and reaching the surface, as island after island slowly disappeared. Thus areas of immense extent in the central and most profound parts of the great oceans, might become interspersed with coral-islets, none of which would rise to a greater height than that attained by detritus heaped up by the sea, and nevertheless they might all have been formed by corals, which absolutely required for their growth a solid foundation within a few fathoms of the surface.

It would be out of place here to do more than allude to the many facts, showing that the supposition of a gradual subsidence over large areas is by no means improbable. We have the clearest proof that a movement of this kind is possible, in the upright trees buried under the strata many thousand feet in thickness; we have also every reason for believing that there are now large areas gradually sinking, in the same manner as others are rising. And when we consider how many parts of the surface of the globe have been elevated within recent geological periods, we must admit that there have been subsidences on a corresponding scale, for otherwise the whole globe would have swollen. It is very remarkable that Mr. Lyell ("Principles of Geology," sixth edition, volume iii., page 386.), even in the first edition of his "Principles of Geology," inferred that the amount of subsidence in the Pacific must have exceeded that of elevation, from the area of land being very small relatively to the agents there tending to form it, namely, the growth of coral and volcanic action. But it will be asked, are there any direct proofs of a subsiding movement in those areas, in which subsidence will explain a phenomenon otherwise inexplicable? This, however, can hardly be expected, for it must ever be most difficult, excepting in countries long civilised, to detect a movement, the tendency of which is to conceal the part affected. In barbarous and semi-civilised nations how long might not a slow movement, even of elevation such as that now affecting Scandinavia, have escaped attention!

Mr. Williams (Williams's "Narrative of Missionary Enterprise," page 31.) insists strongly that the traditions of the natives, which he has taken much pains in collecting, do not indicate the appearance of any new islands: but on the theory of a gradual subsidence, all that would be apparent would be, the water sometimes encroaching slowly on the land, and the land again recovering by the accumulation of detritus its former extent, and perhaps sometimes the conversion of an atoll with coral islets on it, into a bare or into a sunken annular reef. Such changes would naturally take place at the periods when the sea rose above its usual limits, during a gale of more than ordinary strength; and the effects of the two causes would be hardly distinguishable. In Kotzebue's "Voyage" there are accounts of islands, both in the Caroline and Marshall Archipelagoes, which have been partly washed away during hurricanes; and Kadu, the native who was on board one of the Russian vessels, said "he saw the sea at Radack rise to the feet of the cocoa-nut trees; but it was conjured in time." (Kotzebue's "First Voyage," volume iii., page 168.) A storm lately entirely swept away two of the Caroline islands, and converted them into shoals; it partly, also, destroyed two other islands. (M. Desmoulins in "Comptes Rendus," 1840, page 837.) According to a tradition which was communicated to Captain Fitzroy, it is believed in the Low Archipelago, that the arrival of the first ship caused a great inundation, which destroyed many lives. Mr. Stutchbury relates, that in 1825, the western side of Chain Atoll, in the same group, was completely devastated by a hurricane, and not less than 300 lives lost: "in this instance it was evident, even to the natives, that the hurricane alone was not sufficient to account for the violent agitation of the ocean." ("West of England Journal", No. I., page 35.) That considerable changes have taken place recently in some of the atolls in the Low Archipelago, appears certain from the case already given of Matilda Island: with respect to Whitsunday and Gloucester Islands in this same group, we must either attribute great inaccuracy to their discoverer, the famous circumnavigator Wallis, or believe that they have undergone a considerable change in the period of fifty-nine years, between his voyage and that of Captain Beechey's. Whitsunday Island is described by Wallis as "about four miles long, and three wide," now it is only one mile and a half long. The appearance of Gloucester Island, in Captain Beechey's words (Beechey's "Voyage to the Pacific," chapter vii., and Wallis's "Voyage in the 'Dolphin'," chapter iv.), has been accurately described by its discoverer, but its present form and extent differ materially." Blenheim reef, in the Chagos group, consists of a water-washed annular reef, thirteen miles in circumference, surrounding a lagoon ten fathoms deep: on its surface there were a few worn patches of conglomerate coral-rock, of about the size of hovels; and these Captain Moresby considered as being, without doubt, the last remnants of islets; so that here an atoll has been converted into an atoll-formed reef. The inhabitants of the Maldiva Archipelago, as long ago as 1605, declared, "that the high tides and violent currents were diminishing the number of the islands" (See an extract from Pyrard's Voyage in Captain Owen's paper on the Maldiva Archipelago, in the "Geographical Journal", volume ii., page 84.): and I have already shown, on the authority of Captain Moresby, that the work of destruction is still in progress; but that on the other hand the first formation of some islets is known to the present inhabitants. In such cases, it would be exceedingly difficult to detect a gradual subsidence of the foundation, on which these mutable structures rest.

Some of the archipelagoes of low coral-islands are subject to earthquakes: Captain Moresby informs me that they are frequent, though not very strong, in the Chagos group, which occupies a very central position in the Indian Ocean, and is far from any land not of coral formation. One of the islands in this group was formerly covered by a bed of mould, which, after an earthquake, disappeared, and was believed by the residents to have been washed by the rain through the broken masses of underlying rock; the island was thus rendered unproductive. Chamisso (See Chamisso, in Kotzebue's "First Voyage," volume iii., pages 182 and 136.) states, that earthquakes are felt in the Marshall atolls, which are far from any high land, and likewise in the islands of the Caroline Archipelago. On one of the latter, namely Oulleay atoll, Admiral Lutke, as he had the kindness to inform me, observed several straight fissures about a foot in width, running for some hundred yards obliquely across the whole width of the reef. Fissures indicate a stretching of the earth's crust, and, therefore, probably changes in its level; but these coral-islands, which have been shaken and fissured, certainly have not been elevated, and, therefore, probably they have subsided. In the chapter on Keeling atoll, I attempted to show by direct evidence, that the island underwent a movement of subsidence, during the earthquakes lately felt there.

The facts stand thus;—there are many large tracts of ocean, without any high land, interspersed with reefs and islets, formed by the growth of those kinds of corals, which cannot live at great depths; and the existence of these reefs and low islets, in such numbers and at such distant points, is quite inexplicable, excepting on the theory, that the bases on which the reefs first became attached, slowly and successively sank beneath the level of the sea, whilst the corals continued to grow upwards. No positive facts are opposed to this view, and some general considerations render it probable. There is evidence of change in form, whether or not from subsidence, on some of these coral-islands; and there is evidence of subterranean disturbances beneath them. Will then the theory, to which we have thus been led, solve the curious problem,—what has given to each class of reef its peculiar form?

(PLATE: WOODCUT NO. 4.

AA—Outer edge of the reef at the level of the sea.

BB—Shores of the island.

A'A'—Outer edge of the reef, after its upward growth during a period of subsidence.

CC—The lagoon-channel between the reef and the shores of the now encircled land.

B'B'—The shores of the encircled island.

N.B.—In this, and the following woodcut, the subsidence of the land could only be represented by an apparent rise in the level of the sea.

PLATE: WOODCUT NO. 5.

A'A'—Outer edges of the barrier-reef at the level of the sea. The cocoa-nut trees represent coral-islets formed on the reef.

CC—The lagoon-channel.

B'B'—The shores of the island, generally formed of low alluvial land and of coral detritus from the lagoon-channel.

A"A"—The outer edges of the reef now forming an atoll.

C'—The lagoon of the newly formed atoll. According to the scale, the depth of the lagoon and of the lagoon-channel is exaggerated.)

Let us in imagination place within one of the subsiding areas, an island surrounded by a "fringing-reef,"—that kind, which alone offers no difficulty in the explanation of its origin. Let the unbroken lines and the oblique shading in the woodcut (No. 4) represent a vertical section through such an island; and the horizontal shading will represent the section of the reef. Now, as the island sinks down, either a few feet at a time or quite insensibly, we may safely infer from what we know of the conditions favourable to the growth of coral, that the living masses bathed by the surf on the margin of the reef, will soon regain the surface. The water, however, will encroach, little by little, on the shore, the island becoming lower and smaller, and the space between the edge of the reef and the beach proportionately broader. A section of the reef and island in this state, after a subsidence of several hundred feet, is given by the dotted lines: coral-islets are supposed to have been formed on the new reef, and a ship is anchored in the lagoon-channel. This section is in every respect that of an encircling barrier-reef; it is, in fact, a section taken (The section has been made from the chart given in the "Atlas of the Voyage of the 'Coquille'." The scale is .57 of an inch to a mile. The height of the island, according to M. Lesson, is 4,026 feet. The deepest part of the lagoon-channel is 162 feet; its depth is exaggerated in the woodcut for the sake of clearness.) east and west through the highest point of the encircled island of Bolabola; of which a plan is given in Plate I., Figure 5. The same section is more clearly shown in the following woodcut (No. 5) by the unbroken lines. The width of the reef, and its slope, both on the outer and inner side, will have been determined by the growing powers of the coral, under the conditions (for instance the force of the breakers and of the currents) to which it has been exposed; and the lagoon-channel will be deeper or shallower, in proportion to the growth of the delicately branched corals within the reef, and to the accumulation of sediment, relatively, also, to the rate of subsidence and the length of the intervening stationary periods.

It is evident in this section, that a line drawn perpendicularly down from the outer edge of the new reef to the foundation of solid rock, exceeds by as many feet as there have been feet of subsidence, that small limit of depth at which the effective polypifers can live—the corals having grown up, as the whole sank down, from a basis formed of other corals and their consolidated fragments. Thus the difficulty on this head, which before seemed so great, disappears.

As the space between the reef and the subsiding shore continued to increase in breadth and depth, and as the injurious effects of the sediment and fresh water borne down from the land were consequently lessened, the greater number of the channels, with which the reef in its fringing state must have been breached, especially those which fronted the smaller streams, will have become choked up with the growth of coral: on the windward side of the reef, where the coral grows most vigorously, the breaches will probably have first been closed. In barrier-reefs, therefore, the breaches kept open by draining the tidal waters of the lagoon-channel, will generally be placed on the leeward side, and they will still face the mouths of the larger streams, although removed beyond the influence of their sediment and fresh water;—and this, it has been shown, is commonly the case.

Referring to the diagram shown above, in which the newly formed barrier-reef is represented by unbroken lines, instead of by dots as in the former woodcut, let the work of subsidence go on, and the doubly pointed hill will form two small islands (or more, according to the number of the hills) included within one annular reef. Let the island continue subsiding, and the coral-reef will continue growing up on its own foundation, whilst the water gains inch by inch on the land, until the last and highest pinnacle is covered, and there remains a perfect atoll. A vertical section of this atoll is shown in the woodcut by the dotted lines;—a ship is anchored in its lagoon, but islets are not supposed yet to have been formed on the reef. The depth of the lagoon and the width and slope of the reef, will depend on the circumstances just referred to under barrier-reefs. Any further subsidence will produce no change in the atoll, except perhaps a diminution in its size, from the reef not growing vertically upwards; but should the currents of the sea act violently upon it, and should the corals perish on part or on the whole of its margin, changes would result during subsidence which will be presently noticed. I may here observe, that a bank either of rock or of hardened sediment, level with the surface of the sea, and fringed with living coral, would (if not so small as to allow the central space to be quickly filled up with detritus) by subsidence be converted immediately into an atoll, without passing, as in the case of a reef fringing the shore of an island, through the intermediate form of a barrier-reef. If such a bank lay a few fathoms submerged, the simple growth of the coral (as remarked in the third chapter) without the aid of subsidence, would produce a structure scarcely to be distinguished from a true atoll; for in all cases the corals on the outer margin of a reef, from having space and being freely exposed to the open sea, will grow vigorously and tend to form a continuous ring whilst the growth of the less massive kinds on the central expanse, will be checked by the sediment formed there, and by that washed inwards by the breakers; and as the space becomes shallower, their growth will, also, be checked by the impurities of the water, and probably by the small amount of food brought by the enfeebled currents, in proportion to the surface of living reefs studded with innumerable craving mouths: the subsidence of a reef based on a bank of this kind, would give depth to its central expanse or lagoon, steepness to its flanks, and through the free growth of the coral, symmetry to its outline:—I may here repeat that the larger groups of atolls in the Pacific and Indian Oceans cannot be supposed to be founded on banks of this nature.

If, instead of the island in the diagram, the shore of a continent fringed by a reef had subsided, a great barrier-reef, like that on the north-east coast of Australia, would have necessarily resulted; and it would have been separated from the main land by a deep-water channel, broad in proportion to the amount of subsidence, and to the less or greater inclination of the neighbouring coast-line. The effect of the continued subsidence of a great barrier-reef of this kind, and its probable conversion into a chain of separate atolls, will be noticed, when we discuss the apparent progressive disseverment of the larger Maldiva atolls.

We now are able to perceive that the close similarity in form, dimensions, structure, and relative position (which latter point will hereafter be more fully noticed) between fringing and encircling barrier-reefs, and between these latter and atolls, is the necessary result of the transformation, during subsidence of the one class into the other. On this view, the three classes of reefs ought to graduate into each other. Reefs having intermediate character between those of the fringing and barrier classes do exist; for instance, on the south-west coast of Madagascar, a reef extends for several miles, within which there is a broad channel from seven to eight fathoms deep, but the sea does not deepen abruptly outside the reef. Such cases, however, are open to some doubts, for an old fringing-reef, which had extended itself a little on a basis of its own formation, would hardly be distinguishable from a barrier-reef, produced by a small amount of subsidence, and with its lagoon-channel nearly filled up with sediment during a long stationary period. Between barrier-reefs, encircling either one lofty island or several small low ones, and atolls including a mere expanse of water, a striking series can be shown: in proof of this, I need only refer to the first plate in this volume, which speaks more plainly to the eye, than any description could to the ear. The authorities from which the charts have been engraved, together with some remarks on them and descriptive of the plates, are given above. At New Caledonia (Plate II., Figure 5.) the barrier-reefs extend for 150 miles on each side of the submarine prolongation of the island; and at their northern extremity they appear broken up and converted into a vast atoll-formed reef, supporting a few low coral-islets: we may imagine that we here see the effects of subsidence actually in progress, the water always encroaching on the northern end of the island, towards which the mountains slope down, and the reefs steadily building up their massive fabrics in the lines of their ancient growth.

We have as yet only considered the origin of barrier-reefs and atolls in their simplest form; but there remain some peculiarities in structure and some special cases, described in the two first chapters, to be accounted for by our theory. These consist—in the inclined ledge terminated by a wall, and sometimes succeeded by a second ledge with a wall, round the shores of certain lagoons and lagoon-channels; a structure which cannot, as I endeavoured to show, be explained by the simple growing powers of the corals,—in the ring or basin-like forms of the central reefs, as well as of the separate marginal portions of the northern Maldiva atolls,—in the submerged condition of the whole, or of parts of certain barrier and atoll-formed reefs; where only a part is submerged, this being generally to leeward,—in the apparent progressive disseverment of some of the Maldiva atolls,—in the existence of irregularly formed atolls, some being tied together by linear reefs, and others with spurs projecting from them,—and, lastly, in the structure and origin of the Great Chagos Bank.

STEP-FORMED LEDGES ROUND CERTAIN LAGOONS.

If we suppose an atoll to subside at an extremely slow rate, it is difficult to follow out the complex results. The living corals would grow up on the outer margin; and likewise probably in the gullies and deeper parts of the bare surface of the annular reef; the water would encroach on the islets, but the accumulation of fresh detritus might possibly prevent their entire submergence. After a subsidence of this very slow nature, the surface of the annular reef sloping gently into the lagoon, would probably become united with the irregular reefs and banks of sand, which line the shores of most lagoons. Should, however, the atoll be carried down by a more rapid movement, the whole surface of the annular reef, where there was a foundation of solid matter, would be favourably circumstanced for the fresh growth of coral; but as the corals grew upwards on its exterior margin, and the waves broke heavily on this part, the increase of the massive polypifers on the inner side would be checked from the want of water. Consequently, the exterior parts would first reach the surface, and the new annular reef thus formed on the old one, would have its summit inclined inwards, and be terminated by a subaqueous wall, formed by the upward growth of the coral (before being much checked), from the inner edge of the solid parts of the old reef. The inner portion of the new reef, from not having grown to the surface, would be covered by the waters of the lagoon. Should a subsidence of the same kind be repeated, the corals would again grow up in a wall, from all the solid parts of the resunken reef, and, therefore, not from within the sandy shores of the lagoon; and the inner part of the new annular reef would, from being as before checked in its upward growth, be of less height than the exterior parts, and therefore would not reach the surface of the lagoon. In this case the shores of the lagoon would be surrounded by two inclined ledges, one beneath the other, and both abruptly terminated by subaqueous cliffs. (According to Mr. Couthouy (page 26) the external reef round many atolls descends by a succession of ledges or terraces. He attempts, I doubt whether successfully, to explain this structure somewhat in the same manner as I have attempted, with respect to the internal ledges round the lagoons of some atolls. More facts are wanted regarding the nature both of the interior and exterior step-like ledges: are all the ledges, or only the upper ones, covered with living coral? If they are all covered, are the kinds different on the ledges according to the depth? Do the interior and exterior ledges occur together in the same atolls; if so, what is their total width, and is the intervening surface-reef narrow, etc.?)

THE RING OR BASIN-FORMED REEFS OF THE NORTHERN MALDIVA ATOLLS.

I may first observe, that the reefs within the lagoons of atolls and within lagoon-channels, would, if favourably circumstanced, grow upwards during subsidence in the same manner as the annular rim; and, therefore, we might expect that such lagoon-reefs, when not surrounded and buried by an accumulation of sediment more rapid than the rate of subsidence, would rise abruptly from a greater depth than that at which the efficient polypifers can flourish: we see this well exemplified in the small abruptly-sided reefs, with which the deep lagoons of the Chagos and Southern Maldiva atolls are studded. With respect to the ring or basin-formed reefs of the Northern Maldiva atolls, it is evident, from the perfectly continuous series which exists that the marginal rings, although wider than the exterior or bounding reef of ordinary atolls, are only modified portions of such a reef; it is also evident that the central rings, although wider than the knolls or reefs which commonly occur in lagoons, occupy their place. The ring-like structure has been shown to be contingent on the breaches into the lagoon being broad and numerous, so that all the reefs which are bathed by the waters of the lagoon are placed under nearly the same conditions with the outer coast of an atoll standing in the open sea. Hence the exterior and living margins of these reefs must have been favourably circumstanced for growing outwards, and increasing beyond the usual breadth; and they must likewise have been favourably circumstanced for growing vigorously upwards, during the subsiding movements, to which by our theory the whole archipelago has been subjected; and subsidence with this upward growth of the margins would convert the central space of each little reef into a small lagoon. This, however, could only take place with those reefs, which had increased to a breadth sufficient to prevent their central spaces from being almost immediately filled up with the sand and detritus driven inwards from all sides: hence it is that few reefs, which are less than half a mile in diameter, even in the atolls where the basin-like structure is most strikingly exhibited, include lagoons. This remark, I may add, applies to all coral-reefs wherever found. The basin-formed reefs of the Maldiva Archipelago may, in fact, be briefly described, as small atolls formed during subsidence over the separate portions of large and broken atolls, in the same manner as these latter were formed over the barrier-reefs, which encircled the islands of a large archipelago now wholly submerged.

SUBMERGED AND DEAD REEFS.

In the second section of the first chapter, I have shown that there are in the neighbourhood of atolls, some deeply submerged banks, with level surfaces; that there are others, less deeply but yet wholly submerged, having all the characters of perfect atolls, but consisting merely of dead coral-rock; that there are barrier-reefs and atolls with merely a portion of their reef, generally on the leeward side, submerged; and that such portions either retain their perfect outline, or they appear to be quite effaced, their former place being marked only by a bank, conforming in outline with that part of the reef which remains perfect. These several cases are, I believe, intimately related together, and can be explained by the same means. There, perhaps, exist some submerged reefs, covered with living coral and growing upwards, but to these I do not here refer.

As we see that in those parts of the ocean, where coral-reefs are most abundant, one island is fringed and another neighbouring one is not fringed; as we see in the same archipelago, that all the reefs are more perfect in one part of it than in another, for instance, in the southern half compared with the northern half of the Maldiva Archipelago, and likewise on the outer coasts compared with the inner coasts of the atolls in this same group, which are placed in a double row; as we know that the existence of the innumerable polypifers forming a reef, depends on their sustenance, and that they are preyed on by other organic beings; and, lastly, as we know that some inorganic causes are highly injurious to the growth of coral, it cannot be expected that during the round of change to which earth, air, and water are exposed, the reef-building polypifers should keep alive for perpetuity in any one place; and still less can this be expected, during the progressive subsidences, perhaps at some periods more rapid than at others, to which by our theory these reefs and islands have been subjected and are liable. It is, then, not improbable that the corals should sometimes perish either on the whole or on part of a reef; if on part, the dead portion, after a small amount of subsidence, would still retain its proper outline and position beneath the water. After a more prolonged subsidence, it would probably form, owing to the accumulation of sediment, only the margin of a flat bank, marking the limits of the former lagoon. Such dead portions of reef would generally lie on the leeward side (Mr. Lyell, in the first edition of his "Principles of Geology," offered a somewhat different explanation of this structure. He supposes that there has been subsidence; but he was not aware that the submerged portions of reef were in most cases, if not in all, dead; and he attributes the difference in height in the two sides of most atolls, chiefly to the greater accumulation of detritus to windward than to leeward. But as matter is accumulated only on the backward part of the reef, the front part would remain of the same height on both sides. I may here observe that in most cases (for instance, at Peros Banhos, the Gambier group and the Great Chagos Bank), and I suspect in all cases, the dead and submerged portions do not blend or slope into the living and perfect parts, but are separated from them by an abrupt line. In some instances small patches of living reef rise to the surface from the middle of the submerged and dead parts.), for the impure water and fine sediment would more easily flow out from the lagoon over this side of the reef, where the force of the breakers is less than to windward; and therefore the corals would be less vigorous on this side, and be less able to resist any destroying agent. It is likewise owing to this same cause, that reefs are more frequently breached to leeward by narrow channels, serving as by ship-channels, than to windward. If the corals perished entirely, or on the greater part of the circumference of an atoll, an atoll-shaped bank of dead rock, more or less entirely submerged, would be produced; and further subsidence, together with the accumulation of sediment, would often obliterate its atoll-like structure, and leave only a bank with a level surface.

In the Chagos group of atolls, within an area of 160 miles by 60, there are two atoll-formed banks of dead rock (besides another very imperfect one), entirely submerged; a third, with merely two or three very small pieces of living reef rising to the surface; and a fourth, namely, Peros Banhos (Plate I., Figure 9), with a portion nine miles in length dead and submerged. As by our theory this area has subsided, and as there is nothing improbable in the death, either from changes in the state of the surrounding sea or from the subsidence being great or sudden, of the corals on the whole, or on portions of some of the atolls, the case of the Chagos group presents no difficulty. So far indeed are any of the above-mentioned cases of submerged reefs from being inexplicable, that their occurrence might have been anticipated on our theory, and as fresh atolls are supposed to be in progressive formation by the subsidence of encircling barrier-reefs, a weighty objection, namely that the number of atolls must be increasing infinitely, might even have been raised, if proofs of the occasional destruction and loss of atolls could not have been adduced.

THE DISSEVERMENT OF THE LARGER MALDIVA ATOLLS.

The apparent progressive disseverment in the Maldiva Archipelago of large atolls into smaller ones, is, in many respects, an important consideration, and requires an explanation. The graduated series which marks, as I believe, this process, can be observed only in the northern half of the group, where the atolls have exceedingly imperfect margins, consisting of detached basin-formed reefs. The currents of the sea flow across these atolls, as I am informed by Captain Moresby, with considerable force, and drift the sediment from side to side during the monsoons, transporting much of it seaward; yet the currents sweep with greater force round their flanks. It is historically known that these atolls have long existed in their present state; and we can believe, that even during a very slow subsidence they might thus remain, the central expanse being kept at nearly its original depth by the accumulation of sediment. But in the action of such nicely balanced forces during a progressive subsidence (like that, to which by our theory this archipelago has been subjected), it would be strange if the currents of the sea should never make a direct passage across some one of the atolls, through the many wide breaches in their margins. If this were once effected, a deep-water channel would soon be formed by the removal of the finer sediment, and the check to its further accumulation; and the sides of the channel would be worn into a slope like that on the outer coasts, which are exposed to the same force of the currents. In fact, a channel precisely like that bifurcating one which divides Mahlos Mahdoo (Plate II., Figure 4.), would almost necessarily be formed. The scattered reefs situated near the borders of the new ocean-channel, from being favourably placed for the growth of coral, would, by their extension, tend to produce fresh margins to the dissevered portions; such a tendency is very evident (as may be seen in the large published chart) in the elongated reefs on the borders of the two channels intersecting Mahlos Mahdoo. Such channels would become deeper with continued subsidence, and probably from the reefs not growing up perpendicularly, somewhat broader. In this case, and more especially if the channels had been formed originally of considerable breadth, the dissevered portions would become perfect and distinct atolls, like Ari and Ross atolls (Plate II., Figure 6), or like the two Nillandoo atolls, which must be considered as distinct, although related in form and position, and separated from each other by channels, which though deep have been sounded. Further subsidence would render such channels unfathomable, and the dissevered portions would then resemble Phaleedoo and Moluque atolls, or Mahlos Mahdoo and Horsburgh atolls (Plate II., Figure 4), which are related to each other in no respect except in proximity and position. Hence, on the theory of subsidence, the disseverment of large atolls, which have imperfect margins (for otherwise their disseverment would be scarcely possible), and which are exposed to strong currents, is far from being an improbable event; and the several stages, from close relation to entire isolation in the atolls of the Maldiva Archipelago, are readily explicable.

We might go even further, and assert as not improbable, that the first formation of the Maldiva Archipelago was due to a barrier-reef, of nearly the same dimensions with that of New Caledonia (Plate II., Figure 5), for if, in imagination, we complete the subsidence of that great island, we might anticipate from the present broken condition of the northern portion of the reef, and from the almost entire absence of reefs on the eastern coast, that the barrier-reef after repeated subsidences, would become during its upward growth separated into distinct portions; and these portions would tend to assume an atoll-like structure, from the coral growing with vigour round their entire circumferences, when freely exposed to an open sea. As we have some large islands partly submerged with barrier-reefs marking their former limits, such as New Caledonia, so our theory makes it probable that there should be other large islands wholly submerged; and these, we may now infer, would be surmounted, not by one enormous atoll, but by several large elongated ones, like the atolls in the Maldiva group; and these again, during long periods of subsidence, would sometimes become dissevered into smaller atolls. I may add, that both in the Marshall and Caroline Archipelagoes, there are atolls standing close together, which have an evident relationship in form: we may suppose, in such cases, either that two or more encircled islands originally stood close together, and afforded bases for two or more atolls, or that one atoll has been dissevered. From the position, as well as form, of three atolls in the Caroline Archipelago (the Namourrek and Elato group), which are placed in an irregular circle, I am strongly tempted to believe that they have originated by the process of disseverment. (The same remark is, perhaps, applicable to the islands of Ollap, Fanadik, and Tamatam in the Caroline Archipelago, of which charts are given in the atlas of Duperrey's voyage: a line drawn through the linear reefs and lagoons of these three islands forms a semicircle. Consult also, the atlas of Lutke's voyage; and for the Marshall group that of Kotzebue; for the Gilbert group consult the atlas of Duperrey's voyage. Most of the points here referred to may, however, be seen in Krusenstern's general Atlas of the Pacific.)

IRREGULARLY FORMED ATOLLS.

In the Marshall group, Musquillo atoll consists of two loops united in one point; and Menchikoff atoll is formed of three loops, two of which (as may be seen in Figure 3, Plate II.) are connected by a mere ribbon-shaped reef, and the three together are sixty miles in length. In the Gilbert group some of the atolls have narrow strips of reef, like spurs, projecting from them. There occur also in parts of the open sea, a few linear and straight reefs, standing by themselves; and likewise some few reefs in the form of crescents, with their extremities more or less curled inwards. Now, the upward growth of a barrier-reef which fronted only one side of an island, or one side of an elongated island with its extremities (of which cases exist), would produce after the complete subsidence of the land, mere strips or crescent or hook-formed reefs: if the island thus partially fronted became divided during subsidence into two or more islands, these islands would be united together by linear reefs; and from the further growth of the coral along their shores together with subsidence, reefs of various forms might ultimately be produced, either atolls united together by linear reefs, or atolls with spurs projecting from them. Some, however, of the more simple forms above specified, might, as we have seen, be equally well produced by the coral perishing during subsidence on part of the circumference of an atoll, whilst on the other parts it continued to grow up till it reached the surface.

THE GREAT CHAGOS BANK.

I have already shown that the submerged condition of the Great Chagos Bank (Plate II., Figure 1, with its section Figure 2), and of some other banks in the Chagos group, may in all probability be attributed to the coral having perished before or during the movements of subsidence, to which this whole area by our theory has been subjected. The external rim or upper ledge (shaded in the chart), consists of dead coral-rock thinly covered with sand; it lies at an average depth of between five and eight fathoms, and perfectly resembles in form the annular reef of an atoll. The banks of the second level, the boundaries of which are marked by dotted lines in the chart, lie from about fifteen to twenty fathoms beneath the surface; they are several miles broad, and terminate in a very steep slope round the central expanse. This central expanse I have already described, as consisting of a level muddy flat between thirty and forty fathoms deep. The banks of the second level, might at first sight be thought analogous to the internal step-like ledge of coral-rock which borders the lagoons of some atolls, but their much greater width, and their being formed of sand, are points of essential difference. On the eastern side of the atoll some of the banks are linear and parallel, resembling islets in a great river, and pointed directly towards a great breach on the opposite side of the atoll; these are best seen in the large published chart. I inferred from this circumstance, that strong currents sometimes set directly across this vast bank; and I have since heard from Captain Moresby that this is the case. I observed, also, that the channels or breaches through the rim, were all of the same depth as the central lagoon-like space into which they lead; whereas the channels into the other atolls of the Chagos group, and as I believe into most other large atolls, are not nearly as deep as their lagoons: for instance at Peros Banhos, the channels are only of the same depth, namely between ten and twenty fathoms, as the bottom of the lagoon for a space about a mile and a half in width round its shores, whilst the central expanse of the lagoon is from thirty-five to forty fathoms deep. Now, if an atoll during a gradual subsidence once became entirely submerged, like the Great Chagos Bank, and therefore no longer exposed to the surf, very little sediment could be formed from it; and consequently the channels leading into the lagoon from not being filled up with drifted sand and coral detritus, would continue increasing in depth, as the whole sank down. In this case, we might expect that the currents of the open sea, instead of any longer sweeping round the submarine flanks, would flow directly through the breaches across the lagoon, removing in their course the finer sediment, and preventing its further accumulation. We should then have the submerged reef forming an external and upper rim of rock, and beneath this portion of the sandy bottom of the old lagoon, intersected by deep-water channels or breaches, and thus formed into separate marginal banks; and these would be cut off by steep slopes, overhanging the central space, worn down by the passage of the oceanic currents.

By these means, I have scarcely any doubt that the Great Chagos Bank has originated,—a structure which at first appeared to me far more anomalous than any I had met with. The process of formation is nearly the same with that, by which Mahlos Mahdoo had been trisected; but in the Chagos Bank the channels of the oceanic currents entering at several different quarters, have united in a central space.

This great atoll-formed bank appears to be in an early stage of disseverment; should the work of subsidence go on, from the submerged and dead condition of the whole reef, and the imperfection of the south-east quarter a mere wreck would probably be left. The Pitt's Bank, situated not far southward, appears to be precisely in this state; it consists of a moderately level, oblong bank of sand, lying from 10 to 20 fathoms beneath the surface, with two sides protected by a narrow ledge of rock which is submerged between 5 and 8 fathoms. A little further south, at about the same distance as the southern rim of the Great Chagos Bank is from the northern rim, there are two other small banks with from 10 to 20 fathoms on them; and not far eastward soundings were struck on a sandy bottom, with between 110 and 145 fathoms. The northern portion with its ledge-like margin, closely resembles any one segment of the Great Chagos Bank, between two of the deep-water channels, and the scattered banks, southward appear to be the last wrecks of less perfect portions.

I have examined with care the charts of the Indian and Pacific Oceans, and have now brought before the reader all the examples, which I have met with, of reefs differing from the type of the class to which they belong; and I think it has been satisfactorily shown, that they are all included in our theory, modified by occasional accidents which might have been anticipated as probable. In this course we have seen, that in the lapse of ages encircling barrier-reefs are occasionally converted into atolls, the name of atoll being properly applicable, at the moment when the last pinnacle of encircled land sinks beneath the surface of the sea. We have, also, seen that large atolls during the progressive subsidence of the areas in which they stand, sometimes become dissevered into smaller ones; at other times, the reef-building polypifers having entirely perished, atolls are converted into atoll-formed banks of dead rock; and these again through further subsidence and the accumulation of sediment modified by the force of the oceanic currents, pass into level banks with scarcely any distinguishing character. Thus may the history of an atoll be followed from its first origin, through the occasional accidents of its existence, to its destruction and final obliteration.

OBJECTIONS TO THE THEORY OF THE FORMATION OF ATOLLS AND BARRIER-REEFS.

The vast amount of subsidence, both horizontally or in area, and vertically or in depth, necessary to have submerged every mountain, even the highest, throughout the immense spaces of ocean interspersed with atolls, will probably strike most people as a formidable objection to my theory. But as continents, as large as the spaces supposed to have subsided, have been raised above the level of the sea,—as whole regions are now rising, for instance, in Scandinavia and South America,—and as no reason can be assigned, why subsidences should not have occurred in some parts of the earth's crust on as great a scale both in extent and amount as those of elevation, objections of this nature strike me as of little force. The remarkable point is that movements to such an extent should have taken place within a period, during which the polypifers have continued adding matter on and above the same reefs. Another and less obvious objection to the theory will perhaps be advanced from the circumstance, of the lagoons within atolls and within barrier-reefs never having become in any one instance during prolonged subsidences of a greater depth than sixty fathoms, and seldom more than forty fathoms; but we already admit, if the theory be worth considering, that the rate of subsidence has not exceeded that of the upward growth of the coral on the exterior margin; we are, therefore, only further required to admit, that the subsidence has not exceeded in rate the filling up of the interior spaces by the growth of the corals living there, and by the accumulation of sediment. As this filling up must take place very slowly within barrier-reefs lying far from the land, and within atolls which are of large dimensions and which have open lagoons with very few reefs, we are led to conclude that the subsidence thus counter-balanced, must have been slow in an extraordinary degree; a conclusion which accords with our only means, namely, with what is known of the rate and manner of recent elevatory movements, of judging by analogy what is the probable rate of subsidence.

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