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American Addresses, with a Lecture on the Study of Biology
by Tomas Henry Huxley
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These are the Pterosauria, or Pterodactyles, the remains of which are met with throughout the series of Mesozoic rocks, from the lias to the chalk, and some of which attained a great size, their wings having a span of eighteen or twenty feet. These animals, in the form and proportions of the head and neck relatively to the body, and in the fact that the ends of the jaws were often, if not always, more or less extensively ensheathed in horny beaks, remind us of birds. Moreover, their bones contained air cavities, rendering them specifically lighter, as is the case in most birds. The breast-bone was large and keeled, as in most birds and in bats, and the shoulder girdle is strikingly similar to that of ordinary birds. But, it seems to me, that the special resemblance of pterodactyles to birds ends here, unless I may add the entire absence of teeth which characterizes the great pterodactyles (Pteranodon), discovered by Professor Marsh. All other known pterodactyles have teeth lodged in sockets. In the vertebral column and the hind limbs there are no special resemblances to birds, and when we turn to the wings they are found to be constructed on a totally different principle from those of birds.

There are four fingers. These four fingers are large, and three of them, those which answer to the thumb and two following fingers in my hand—are terminated by claws, while the fourth is enormously prolonged and converted into a great jointed style. You see at once, from what I have stated about a bird's wing, that there could be nothing less like a bird's wing than this is. It concluded by general reasoning that this finger had the office of supporting a web which extended between it and the body. An existing specimen proves that such was really the case, and that the pterodactyles were devoid of feathers, but that the fingers supported a vast web like that of a bat's wing; in fact, there can be no doubt that this ancient reptile flew after the fashion of a bat.

Thus though the pterodactyle is a reptile which has become modified in such a manner as to enable it to fly, and therefore, as might be expected, presents some points of resemblance to other animals which fly; it has, so to speak, gone off the line which leads directly from reptiles to birds, and has become disqualified for the changes which lead to the characteristic organization of the latter class. Therefore, viewed in relation to the classes of reptiles and birds, the pterodactyles appear to me to be, in a limited sense, intercalary forms; but they are not even approximately linear, in the sense of exemplifying those modifications of structure through which the passage from the reptile to the bird took place.



LECTURE III.

THE DEMONSTRATIVE EVIDENCE OF EVOLUTION.

The occurrence of historical facts is said to be demonstrated, when the evidence that they happened is of such a character as to render the assumption that they did not happen in the highest degree improbable; and the question I now have to deal with is, whether evidence in favour of the evolution of animals of this degree of cogency is, or is not, obtainable from the record of the succession of living forms which is presented to us by fossil remains.

Those who have attended to the progress of palaeontology are aware that evidence of the character which I have defined has been produced in considerable and continually-increasing quantity during the last few years. Indeed, the amount and the satisfactory nature of that evidence are somewhat surprising, when we consider the conditions under which alone we can hope to obtain it.

It is obviously useless to seek for such evidence except in localities in which the physical conditions have been such as to permit of the deposit of an unbroken, or but rarely interrupted, series of strata through a long period of time; in which the group of animals to be investigated has existed in such abundance as to furnish the requisite supply of remains; and in which, finally, the materials composing the strata are such as to ensure the preservation of these remains in a tolerably perfect and undisturbed state.

It so happens that the case which, at present, most nearly fulfils all these conditions is that of the series of extinct animals which culminates in the Horses; by which term I mean to denote not merely the domestic animals with which we are all so well acquainted, but their allies, the ass, zebra, quagga, and the like. In short, I use "horses" as the equivalent of the technical name Equidae, which is applied to the whole group of existing equine animals.

The horse is in many ways a remarkable animal; not least so in the fact that it presents us with an example of one of the most perfect pieces of machinery in the living world. In truth, among the works of human ingenuity it cannot be said that there is any locomotive so perfectly adapted to its purposes, doing so much work with so small a quantity of fuel, as this machine of nature's manufacture—the horse. And, as a necessary consequence of any sort of perfection, of mechanical perfection as of others, you find that the horse is a beautiful creature, one of the most beautiful of all land-animals. Look at the perfect balance of its form, and the rhythm and force of its action. The locomotive machinery is, as you are aware, resident in its slender fore and hind limbs; they are flexible and elastic levers, capable of being moved by very powerful muscles; and, in order to supply the engines which work these levers with the force which they expend, the horse is provided with a very perfect apparatus for grinding its food and extracting therefrom the requisite fuel.

Without attempting to take you very far into the region of osteological detail, I must nevertheless trouble you with some statements respecting the anatomical structure of the horse; and, more especially, will it be needful to obtain a general conception of the structure of its fore and hind limbs, and of its teeth. But I shall only touch upon those points which are absolutely essential to our inquiry.

Let us turn in the first place to the fore-limb. In most quadrupeds, as in ourselves, the fore-arm contains distinct bones called the radius and the ulna. The corresponding region in the Horse seem at first to possess but one bone. Careful observation, however, enables us to distinguish in this bone a part which clearly answers to the upper end of the ulna. This is closely united with the chief mass of the bone which represents the radius, and runs out into a slender shaft which may be traced for some distance downwards upon the back of the radius, and then in most cases thins out and vanishes. It takes still more trouble to make sure of what is nevertheless the fact, that a small part of the lower end of the bone of the horse's fore-arm, which is only distinct in a very young foal, is really the lower extremity of the ulna.

What is commonly called the knee of a horse is its wrist. The "cannon bone" answers to the middle bone of the five metacarpal bones, which support the palm of the hand in ourselves. The "pastern," "coronary," and "coffin" bones of veterinarians answer to the joints of our middle fingers, while the hoof is simply a greatly enlarged and thickened nail. But if what lies below the horse's "knee" thus corresponds to the middle finger in ourselves, what has become of the four other fingers or digits? We find in the places of the second and fourth digits only two slender splint-like bones, about two-thirds as long as the cannon bone, which gradually taper to their lower ends and bear no finger joints, or, as they are termed, phalanges. Sometimes, small bony or gristly nodules are to be found at the bases of these two metacarpal splints, and it is probable that these represent rudiments of the first and fifth toes. Thus, the part of the horse's skeleton, which corresponds with that of the human hand, contains one overgrown middle digit, and at least two imperfect lateral digits; and these answer, respectively, to the third, the second, and the fourth fingers in man.

Corresponding modifications are found in the hind limb. In ourselves, and in most quadrupeds, the leg contains two distinct bones, a large bone, the tibia, and a smaller and more slender bone, the fibula. But, in the horse, the fibula seems, at first, to be reduced to its upper end; a short slender bone united with the tibia, and ending in a point below, occupying its place. Examination of the lower end of a young foal's shin-bone, however, shows a distinct portion of osseous matter, which is the lower end of the fibula; so that the, apparently single, lower end of the shin-bone is really made up of the coalesced ends of the tibia and fibula, just as the, apparently single, lower end of the fore-arm bone is composed of the coalesced radius and ulna.

The heel of the horse is the part commonly known as the hock. The hinder cannon bone answers to the middle metatarsal bone of the human foot, the pastern, coronary, and coffin bones, to the middle toe bones; the hind hoof to the nail; as in the fore-foot. And, as in the fore-foot, there are merely two splints to represent the second and the fourth toes. Sometimes a rudiment of a fifth toe appears to be traceable.

The teeth of a horse are not less peculiar than its limbs. The living engine, like all others, must be well stoked if it is to do its work; and the horse, if it is to make good its wear and tear, and to exert the enormous amount of force required for its propulsion, must be well and rapidly fed. To this end, good cutting instruments and powerful and lasting crushers are needful. Accordingly, the twelve cutting teeth of a horse are close-set and concentrated in the fore part of its mouth, like so many adzes or chisels. The grinders or molars are large, and have an extremely complicated structure, being composed of a number of different substances of unequal hardness. The consequence of this is that they wear away at different rates; and, hence, the surface of each grinder is always as uneven as that of a good millstone.

I have said that the structure of the grinding teeth is very complicated, the harder and the softer parts being, as it were, interlaced with one another. The result of this is that, as the tooth wears, the crown presents a peculiar pattern, the nature of which is not very easily deciphered at first; but which it is important we should understand clearly. Each grinding tooth of the upper jaw has an outer wall so shaped that, on the worn crown, it exhibits the form of two crescents, one in front and one behind, with their concave sides turned outwards. From the inner side of the front crescent, a crescentic front ridge passes inwards and backwards, and its inner face enlarges into a strong longitudinal fold or pillar. From the front part of the hinder crescent, a back ridge takes a like direction, and also has its pillar.

The deep interspaces or valleys between these ridges and the outer wall are filled by bony substance, which is called cement, and coats the whole tooth.

The pattern of the worn face of each grinding tooth of the lower jaw is quite different. It appears to be formed of two crescent-shaped ridges, the convexities of which are turned outwards. The free extremity of each crescent has a pillar, and there is a large double pillar where the two crescents meet. The whole structure is, as it were, imbedded in cement, which fills up the valleys, as in the upper grinders.

If the grinding faces of an upper and of a lower molar of the same side are applied together, it will be seen that the apposed ridges are nowhere parallel, but that they frequently cross; and that thus, in the act of mastication, a hard surface in the one is constantly applied to a soft surface in the other, and vice versa. They thus constitute a grinding apparatus of great efficiency, and one which is repaired as fast as it wears, owing to the long-continued growth of the teeth.

Some other peculiarities of the dentition of the horse must be noticed, as they bear upon what I shall have to say by and by. Thus the crowns of the cutting teeth have a peculiar deep pit, which gives rise to the well-known "mark" of the horse. There is a large space between the outer incisors and the front grinder. In this space the adult male horse presents, near the incisors on each side, above and below, a canine or "tush," which is commonly absent in mares. In a young horse, moreover, there is not unfrequently to be seen in front of the first grinder, a very small tooth, which soon falls out. If this small tooth be counted as one, it will be found that there are seven teeth behind the canine on each side; namely, the small tooth in question, and the six great grinders, among which, by an unusual peculiarity, the foremost tooth is rather larger than those which follow it.

I have now enumerated those characteristic structures of the horse which are of most importance for the purpose we have in view.

To any one who is acquainted with the morphology of vertebrated animals, they show that the horse deviates widely from the general structure of mammals; and that the horse type is, in many respects, an extreme modification of the general mammalian plan. The least modified mammals, in fact, have the radius and ulna, the tibia and fibula, distinct and separate. They have five distinct and complete digits on each foot, and no one of these digits is very much larger than the rest. Moreover, in the least modified mammals, the total number of the teeth is very generally forty-four, while in horses, the usual number is forty, and in the absence of the canines, it may be reduced to thirty-six; the incisor teeth are devoid of the fold seen in those of the horse: the grinders regularly diminish in size from the middle of the series to its front end; while their crowns are short, early attain their full length, and exhibit simple ridges or tubercles, in place of the complex foldings of the horse's grinders.

Hence the general principles of the hypothesis of evolution lead to the conclusion that the horse must have been derived from some quadruped which possessed five complete digits on each foot; which had the bones of the fore-arm and of the leg complete and separate; and which possessed forty-four teeth, among which the crowns of the incisors and grinders had a simple structure; while the latter gradually increased in size from before backwards, at any rate in the anterior part of the series, and had short crowns.

And if the horse has been thus evolved, and the remains of the different stages of its evolution have been preserved, they ought to present us with a series of forms in which the number of the digits becomes reduced; the bones of the fore-arm and leg gradually take on the equine condition; and the form and arrangement of the teeth successively approximate to those which obtain in existing horses.

Let us turn to the facts, and see how far they fulfil these requirements of the doctrine of evolution.

In Europe abundant remains of horses are found in the Quaternary and later Tertiary strata as far as the Pliocene formation. But these horses, which are so common in the cave-deposits and in the gravels of Europe, are in all essential respects like existing horses. And that is true of all the horses of the latter part of the Pliocene epoch. But, in deposits which belong to the earlier Pliocene and later Miocene epochs, and which occur in Britain, in France, in Germany, in Greece, in India, we find animals which are extremely like horses—which, in fact, are so similar to horses, that you may follow descriptions given in works upon the anatomy of the horse upon the skeletons of these animals—but which differ in some important particulars. For example, the structure of their fore and hind limbs is somewhat different. The bones which, in the horse, are represented by two splints, imperfect below, are as long as the middle metacarpal and metatarsal bones; and, attached to the extremity of each, is a digit with three joints of the same general character as those of the middle digit, only very much smaller. These small digits are so disposed that they could have had but very little functional importance, and they must have been rather of the nature of the dew-claws, such as are to be found in many ruminant animals. The Hipparion, as the extinct European three-toed horse is called, in fact, presents a foot similar to that of the American Protohippus (Fig. 9), except that, in the Hipparion, the smaller digits are situated farther back, and are of smaller proportional size, than in the Protohippus.

The ulna is slightly more distinct than in the horse; and the whole length of it, as a very slender shaft, intimately united with the radius, is completely traceable. The fibula appears to be in the same condition as in the horse. The teeth of the Hipparion are essentially similar to those of the horse, but the pattern of the grinders is in some respects a little more complex, and there is a depression on the face of the skull in front of the orbit, which is not seen in existing horses.

In the earlier Miocene, and perhaps the later Eocene deposits of some parts of Europe, another extinct animal has been discovered, which Cuvier, who first described some fragments of it, considered to be a Palaeotherium. But as further discoveries threw new light upon its structure, it was recognised as a distinct genus, under the name of Anchitherium.

In its general characters, the skeleton of Anchitherium is very similar to that of the horse. In fact, Lartet and De Blainville called it Palaeotherium equinum or hippoides; and De Christol, in 1847, said that it differed from Hipparion in little more than the characters of its teeth, and gave it the name of Hipparitherium. Each foot possesses three complete toes; while the lateral toes are much larger in proportion to the middle toe than in Hipparion, and doubtless rested on the ground in ordinary locomotion.

The ulna is complete and quite distinct from the radius, though firmly united with the latter. The fibula seems also to have been complete. Its lower end, though intimately united with that of the tibia, is clearly marked off from the latter bone.

There are forty-four teeth. The incisors have no strong pit. The canines seem to have been well developed in both sexes. The first of the seven grinders, which, as I have said, is frequently absent, and, when it does exist, is small in the horse, is a good-sized and permanent tooth, while the grinder which follows it is but little larger than the hinder ones. The crowns of the grinders are short, and though the fundamental pattern of the horse-tooth is discernible, the front and back ridges are less curved, the accessory pillars are wanting, and the valleys, much shallower, are not filled up with cement.

Seven years ago, when I happened to be looking critically into the bearing of palaeontological facts upon the doctrine of evolution, it appeared to me that the Anchitherium, the Hipparion, and the modern horses, constitute a series in which the modifications of structure coincide with the order of chronological occurrence, in the manner in which they must coincide, if the modern horses really are the result of the gradual metamorphosis, in the course of the Tertiary epoch, of a less specialised ancestral form. And I found by correspondence with the late eminent French anatomist and palaeontologist, M. Lartet, that he had arrived at the same conclusion from the same data.

That the Anchitherium type had become metamorphosed into the Hipparion type, and the latter into the Equine type, in the course of that period of time which is represented by the latter half of the Tertiary deposits, seemed to me to be the only explanation of the facts for which there was even a shadow of probability.[2]

And, hence, I have ever since held that these facts afford evidence of the occurrence of evolution, which, in the sense already defined, may be termed demonstrative.

All who have occupied themselves with the structure of Anchitherium, from Cuvier onwards, have acknowledged its many points of likeness to a well-known genus of extinct Eocene mammals, Palaeotherium. Indeed, as we have seen, Cuvier regarded his remains of Anchitherium as those of a species of Palaeotherium. Hence, in attempting to trace the pedigree of the horse beyond the Miocene epoch and the Anchitheroid form, I naturally sought among the various species of Palaeotheroid animals for its nearest ally, and I was led to conclude that the Palaeotherium minus (Plagiolophus) represented the next step more nearly than any form then known.

I think that this opinion was fully justifiable; but the progress of investigation has thrown an unexpected light on the question, and has brought us much nearer than could have been anticipated to a knowledge of the true series of the progenitors of the horse.

You are all aware that, when your country was first discovered by Europeans, there were no traces of the existence of the horse in any part of the American Continent. The accounts of the conquest of Mexico dwell upon the astonishment of the natives of that country when they first became acquainted with that astounding phenomenon—a man seated upon a horse. Nevertheless, the investigations of American geologists have proved that the remains of horses occur in the most superficial deposits of both North and South America, just as they do in Europe. Therefore, for some reason or other—no feasible suggestion on that subject, so far as I know, has been made—the horse must have died out on this continent at some period preceding the discovery of America. Of late years there has been discovered in your Western Territories that marvellous accumulation of deposits, admirably adapted for the preservation of organic remains, to which I referred the other evening, and which furnishes us with a consecutive series of records of the fauna of the older half of the Tertiary epoch, for which we have no parallel in Europe. They have yielded fossils in an excellent state of conservation and in unexampled number and variety. The researches of Leidy and others have shown that forms allied to the Hipparion and the Anchitherium are to be found among these remains. But it is only recently that the admirably conceived and most thoroughly and patiently worked-out investigations of Professor Marsh have given us a just idea of the vast fossil wealth, and of the scientific importance, of these deposits. I have had the advantage of glancing over the collections in Yale Museum; and I can truly say that, so far as my knowledge extends, there is no collection from any one region and series of strata comparable, for extent, or for the care with which the remains have been got together, or for their scientific importance, to the series of fossils which he has deposited there. This vast collection has yielded evidence bearing upon the question of the pedigree of the horse of the most striking character. It tends to show that we must look to America, rather than to Europe, for the original seat of the equine series; and that the archaic forms and successive modifications of the horse's ancestry are far better preserved here than in Europe.

Professor Marsh's kindness has enabled me to put before you a diagram, every figure in which is an actual representation of some specimen which is to be seen at Yale at this present time (Fig. 9).



The succession of forms which he has brought together carries us from the top to the bottom of the Tertiaries. Firstly, there is the true horse. Next we have the American Pliocene form of the horse (Pliohippus); in the conformation of its limbs it presents some very slight deviations from the ordinary horse, and the crowns of the grinding teeth are shorter. Then comes the Protohippus, which represents the European Hipparion, having one large digit and two small ones on each foot, and the general characters of the fore-arm and leg to which I have referred. But it is more valuable than the European Hipparion for the reason that it is devoid of some of the peculiarities of that form—peculiarities which tend to show that the European Hipparion is rather a member of a collateral branch, than a form in the direct line of succession. Next, in the backward order in time, is the Miohippus, which corresponds pretty nearly with the Anchitherium of Europe. It presents three complete toes—one large median and two smaller lateral ones; and there is a rudiment of that digit, which answers to the little finger of the human hand.

The European record of the pedigree of the horse stops here; in the American Tertiaries, on the contrary, the series of ancestral equine forms is continued into the Eocene formations. An older Miocene form, termed Mesohippus, has three toes in front, with a large splint-like rudiment representing the little finger; and three toes behind. The radius and ulna, the tibia and the fibula, are distinct, and the short crowned molar teeth are anchitherioid in pattern.

But the most important discovery of all is the Orohippus, which comes from the Eocene formation, and is the oldest member of the equine series, as yet known. Here we find four complete toes on the front-limb, three toes on the hind-limb, a well-developed ulna, a well-developed fibula, and short-crowned grinders of simple pattern.

Thus, thanks to these important researches, it has become evident that, so far as our present knowledge extends, the history of the horse-type is exactly and precisely that which could have been predicted from a knowledge of the principles of evolution. And the knowledge we now possess justifies us completely in the anticipation, that when the still lower Eocene deposits, and those which belong to the Cretaceous epoch, have yielded up their remains of ancestral equine animals, we shall find, first, a form with four complete toes and a rudiment of the innermost or first digit in front, with, probably, a rudiment of the fifth digit in the hind foot;[3] while, in still older forms, the series of the digits will be more and more complete, until we come to the five-toed animals, in which, if the doctrine of evolution is well founded, the whole series must have taken its origin.

* * * * *

That is what I mean by demonstrative evidence of evolution. An inductive hypothesis is said to be demonstrated when the facts are shown to be in entire accordance with it. If that is not scientific proof, there are no merely inductive conclusions which can be said to be proved. And the doctrine of evolution, at the present time, rests upon exactly as secure a foundation as the Copernican theory of the motions of the heavenly bodies did at the time of its promulgation. Its logical basis is precisely of the same character—the coincidence of the observed facts with theoretical requirements.

The only way of escape, if it be a way of escape, from the conclusions which I have just indicated, is the supposition that all these different equine forms have been created separately at separate epochs of time; and, I repeat, that of such an hypothesis as this there neither is, nor can be, any scientific evidence; and, assuredly, so far as I know, there is none which is supported, or pretends to be supported, by evidence or authority of any other kind. I can but think that the time will come when such suggestions as these, such obvious attempts to escape the force of demonstration, will be put upon the same footing as the supposition made by some writers, who are, I believe, not completely extinct at present, that fossils are mere simulacra, are no indications of the former existence of the animals to which they seem to belong; but that they are either sports of Nature, or special creations, intended—as I heard suggested the other day—to test our faith.

In fact, the whole evidence is in favour of evolution, and there is none against it. And I say this, although perfectly well aware of the seeming difficulties which have been built up upon what appears to the uninformed to be a solid foundation. I meet constantly with the argument that the doctrine of evolution cannot be well founded, because it requires the lapse of a very vast period of time; the duration of life upon the earth, thus implied, is inconsistent with the conclusions arrived at by the astronomer and the physicist. I may venture to say that I am familiar with those conclusions, inasmuch as some years ago, when President of the Geological Society of London, I took the liberty of criticising them, and of showing in what respects, as it appeared to me, they lacked complete and thorough demonstration. But, putting that point aside, suppose that, as the astronomers, or some of them, and some physical philosophers, tell us, it is impossible that life could have endured upon the earth for as long a period as is required by the doctrine of evolution—supposing that to be proved—I desire to be informed, what is the foundation for the statement that evolution does require so great a time? The biologist knows nothing whatever of the amount of time which may be required for the process of evolution. It is a matter of fact that the equine forms which I have described to you occur, in the order stated, in the Tertiary formations. But I have not the slightest means of guessing whether it took a million of years, or ten millions, or a hundred millions, or a thousand millions of years, to give rise to that series of changes. A biologist has no means of arriving at any conclusion as to the amount of time which may be needed for a certain quantity of organic change. He takes his time from the geologist. The geologist, considering the rate at which deposits are formed and the rate at which denudation goes on upon the surface of the earth, arrives at more or less justifiable conclusions as to the time which is required for the deposit of a certain thickness of rocks; and if he tells me that the Tertiary formations required 500,000,000 years for their deposit, I suppose he has good ground for what he says, and I take that as a measure of the duration of the evolution of the horse from the Orohippus up to its present condition. And, if he is right, undoubtedly evolution is a very slow process, and requires a great deal of time. But suppose, now, that an astronomer or a physicist—for instance, my friend Sir William Thomson—tells me that my geological authority is quite wrong; and that he has weighty evidence to show that life could not possibly have existed upon the surface of the earth 500,000,000 years ago, because the earth would have then been too hot to allow of life, my reply is: "That is not my affair; settle that with the geologist, and when you have come to an agreement among yourselves I will adopt your conclusion." We take our time from the geologists and physicists; and it is monstrous that, having taken our time from the physical philosopher's clock, the physical philosopher should turn round upon us, and say we are too fast or too slow. What we desire to know is, is it a fact that evolution took place? As to the amount of time which evolution may have occupied, we are in the hands of the physicist and the astronomer, whose business it is to deal with those questions.

* * * * *

I have now, ladies and gentlemen, arrived at the conclusion of the task which I set before myself when I undertook to deliver these lectures. My purpose has been, not to enable those among you who have paid no attention to these subjects before, to leave this room in a condition to decide upon the validity or the invalidity of the hypothesis of evolution; but I have desired to put before you the principles upon which all hypotheses respecting the history of Nature must be judged; and furthermore, to make apparent the nature of the evidence and the amount of cogency which is to be expected and may be obtained from it. To this end, I have not hesitated to regard you as genuine students and persons desirous of knowing the truth. I have not shrunk from taking you through long discussions, that I fear may have sometimes tried your patience; and I have inflicted upon you details which were indispensable, but which may well have been wearisome. But I shall rejoice—I shall consider that I have done you the greatest service, which it was in my power to do—if I have thus convinced you that the great question which we have been discussing is not one to be dealt with by rhetorical flourishes, or by loose and superficial talk; but that it requires the keen attention of the trained intellect and the patience of the accurate observer.

When I commenced this series of lectures, I did not think it necessary to preface them with a prologue, such as might be expected from a stranger and a foreigner; for during my brief stay in your country, I have found it very hard to believe that a stranger could be possessed of so many friends, and almost harder that a foreigner could express himself in your language in such a way as to be, to all appearance, so readily intelligible. So far as I can judge, that most intelligent, and, perhaps, I may add, most singularly active and enterprising body, your press reporters, do not seem to have been deterred by my accent from giving the fullest account of everything that I happen to have said.

But the vessel in which I take my departure to-morrow morning is even now ready to slip her moorings; I awake from my delusion that I am other than a stranger and a foreigner. I am ready to go back to my place and country; but, before doing so, let me, by way of epilogue, tender to you my most hearty thanks for the kind and cordial reception which you have accorded to me; and let me thank you still more for that which is the greatest compliment which can be afforded to any person in my position—the continuous and undisturbed attention which you have bestowed upon the long argument which I have had the honour to lay before you.

[1] The absence of any keel on the breast-bone and some other osteological peculiarities, observed by Professor Marsh, however, suggest that Hesperornis may be a modification of a less specialised group of birds than that to which these existing aquatic birds belong.

[2] I use the word "type" because it is highly probable that many forms of Anchitherium-like and Hipparion-like animals existed in the Miocene and Pliocene epochs, just as many species of the horse tribe exist now; and it is highly improbable that the particular species of Anchitherium or Hipparion, which happen to have been discovered, should be precisely those which have formed part of the direct line of the horse's pedigree.

[3] Since this lecture was delivered, Professor Marsh has discovered a new genus of equine mammals (Eohippus) from the lowest Eocene deposits of the West, which corresponds very nearly to this description.—American Journal of Science, November, 1876.



BALTIMORE.

ADDRESS ON UNIVERSITY EDUCATION.[1]

The actual work of the University founded in this city by the well-considered munificence of Johns Hopkins commences to-morrow, and among the many marks of confidence and good-will which have been bestowed upon me in the United States, there is none which I value more highly than that conferred by the authorities of the University when they invited me to deliver an address on such an occasion.

For the event which has brought us together is, in many respects, unique. A vast property is handed over to an administrative body, hampered by no conditions save these;—That the principal shall not be employed in building: that the funds shall be appropriated, in equal proportions, to the promotion of natural knowledge and to the alleviation of the bodily sufferings of mankind; and, finally, that neither political nor ecclesiastical sectarianism shall be permitted to disturb the impartial distribution of the testator's benefactions.

In my experience of life a truth which sounds very much like a paradox has often asserted itself; namely, that a man's worst difficulties begin when he is able to do as he likes. So long as a man is struggling with obstacles he has an excuse for failure or shortcoming; but when fortune removes them all and gives him the power of doing as he thinks best, then comes the time of trial. There is but one right, and the possibilities of wrong are infinite. I doubt not that the trustees of the Johns Hopkins University felt the full force of this truth when they entered on the administration of their trust a year and a half ago; and I can but admire the activity and resolution which have enabled them, aided by the able president whom they have selected, to lay down the great outlines of their plan, and carry it thus far into execution. It is impossible to study that plan without perceiving that great care, forethought, and sagacity, have been bestowed upon it, and that it demands the most respectful consideration. I have been endeavouring to ascertain how far the principles which underlie it are in accordance with those which have been established in my own mind by much and long-continued thought upon educational questions. Permit me to place before you the result of my reflections.

Under one aspect a university is a particular kind of educational institution, and the views which we may take of the proper nature of a university are corollaries from those which we hold respecting education in general. I think it must be admitted that the school should prepare for the university, and that the university should crown the edifice, the foundations of which are laid in the school. University education should not be something distinct from elementary education, but should be the natural outgrowth and development of the latter. Now I have a very clear conviction as to what elementary education ought to be; what it really may be, when properly organised; and what I think it will be, before many years have passed over our heads, in England and in America. Such education should enable an average boy of fifteen or sixteen to read and write his own language with ease and accuracy, and with a sense of literary excellence derived from the study of our classic writers: to have a general acquaintance with the history of his own country and with the great laws of social existence; to have acquired the rudiments of the physical and psychological sciences, and a fair knowledge of elementary arithmetic and geometry. He should have obtained an acquaintance with logic rather by example than by precept; while the acquirement of the elements of music and drawing should have been pleasure rather than work.

It may sound strange to many ears if I venture to maintain the proposition that a young person, educated thus far, has had a liberal, though perhaps not a full, education. But it seems to me that such training as that to which I have referred may be termed liberal, in both the senses in which that word is employed, with perfect accuracy. In the first place, it is liberal in breadth. It extends over the whole ground of things to be known and of faculties to be trained, and it gives equal importance to the two great sides of human activity—art and science. In the second place, it is liberal in the sense of being an education fitted for free men; for men to whom every career is open, and from whom their country may demand that they should be fitted to perform the duties of any career. I cannot too strongly impress upon you the fact that, with such a primary education as this, and with no more than is to be obtained by building strictly upon its lines, a man of ability may become a great writer or speaker, a statesman, a lawyer, a man of science, painter, sculptor, architect, or musician. That even development of all a man's faculties, which is what properly constitutes culture, may be effected by such an education, while it opens the way for the indefinite strengthening of any special capabilities with which he may be gifted.

In a country like this, where most men have to carve out their own fortunes and devote themselves early to the practical affairs of life, comparatively few can hope to pursue their studies up to, still less beyond, the age of manhood. But it is of vital importance to the welfare of the community that those who are relieved from the need of making a livelihood, and still more, those who are stirred by the divine impulses of intellectual thirst or artistic genius, should be enabled to devote themselves to the higher service of their kind, as centres of intelligence, interpreters of nature, or creators of new forms of beauty. And it is the function of a university to furnish such men with the means of becoming that which it is their privilege and duty to be. To this end the university need cover no ground foreign to that occupied by the elementary school. Indeed it cannot; for the elementary instruction which I have referred to embraces all the kinds of real knowledge and mental activity possible to man. The university can add no new departments of knowledge, can offer no new fields of mental activity; but what it can do is to intensify and specialise the instruction in each department. Thus literature and philology, represented in the elementary school by English alone, in the university will extend over the ancient and modern languages. History, which, like charity, best begins at home, but, like charity, should not end there, will ramify into anthropology, archaeology, political history, and geography, with the history of the growth of the human mind and of its products in the shape of philosophy, science, and art. And the university will present to the student libraries, museums of antiquities, collections of coins, and the like, which will efficiently subserve these studies. Instruction in the elements of social economy, a most essential, but hitherto sadly-neglected part of elementary education, will develop in the university into political economy, sociology, and law. Physical science will have its great divisions of physical geography, with geology and astronomy; physics; chemistry and biology; represented not merely by professors and their lectures, but by laboratories, in which the students, under guidance of demonstrators, will work out facts for themselves and come into that direct contact with reality which constitutes the fundamental distinction of scientific education. Mathematics will soar into its highest regions; while the high peaks of philosophy may be scaled by those whose aptitude for abstract thought has been awakened by elementary logic. Finally, schools of pictorial and plastic art, of architecture, and of music, will offer a thorough discipline in the principles and practice of art to those in whom lies nascent the rare faculty of aesthetic representation, or the still rarer powers of creative genius.

The primary school and the university are the alpha and omega of education. Whether institutions intermediate between these (so-called secondary schools) should exist, appears to me to be a question of practical convenience. If such schools are established, the important thing is that they should be true intermediaries between the primary school and the university, keeping on the wide track of general culture, and not sacrificing one branch of knowledge for another.

Such appear to me to be the broad outlines of the relations which the university, regarded as a place of education, ought to bear to the school, but a number of points of detail require some consideration, however briefly and imperfectly I can deal with them. In the first place, there is the important question of the limitations which should be fixed to the entrance into the university; or, what qualifications should be required of those who propose to take advantage of the higher training offered by the university. On the one hand, it is obviously desirable that the time and opportunities of the university should not be wasted in conferring such elementary instruction as can be obtained elsewhere; while, on the other hand, it is no less desirable that the higher instruction of the university should be made accessible to every one who can take advantage of it, although he may not have been able to go through any very extended course of education. My own feeling is distinctly against any absolute and defined preliminary examination, the passing of which shall be an essential condition of admission to the university. I would admit to the university any one who could be reasonably expected to profit by the instruction offered to him; and I should be inclined, on the whole, to test the fitness of the student, not by examination before he enters the university, but at the end of his first term of study. If, on examination in the branches of knowledge to which he has devoted himself, he show himself deficient in industry or in capacity, it will be best for the university and best for himself, to prevent him from pursuing a vocation for which he is obviously unfit. And I hardly know of any other method than this by which his fitness or unfitness can be safely ascertained, though no doubt a good deal may be done, not by formal cut and dried examination, but by judicious questioning, at the outset of his career.

Another very important and difficult practical question is, whether a definite course of study shall be laid down for those who enter the university; whether a curriculum shall be prescribed; or whether the student shall be allowed to range at will among the subjects which are open to him. And this question is inseparably connected with another, namely, the conferring of degrees. It is obviously impossible that any student should pass through the whole of the series of courses of instruction offered by a university. If a degree is to be conferred as a mark of proficiency in knowledge, it must be given on the ground that the candidate is proficient in a certain fraction of those studies; and then will arise the necessity of insuring an equivalency of degrees, so that the course by which a degree is obtained shall mark approximately an equal amount of labour and of acquirements, in all cases. But this equivalency can hardly be secured in any other way than by prescribing a series of definite lines of study. This is a matter which will require grave consideration. The important points to bear in mind, I think, are that there should not be too many subjects in the curriculum, and that the aim should be the attainment of thorough and sound knowledge of each.

One half of the Johns Hopkins bequest is devoted to the establishment of a hospital, and it was the desire of the testator that the university and the hospital should co-operate in the promotion of medical education. The trustees will unquestionably take the best advice that is to be had as to the construction and administration of the hospital. In respect to the former point, they will doubtless remember that a hospital may be so arranged as to kill more than it cures; and, in regard to the latter, that a hospital may spread the spirit of pauperism among the well-to-do, as well as relieve the sufferings of the destitute. It is not for me to speak on these topics—rather let me confine myself to the one matter on which my experience as a student of medicine, and an examiner of long standing, who has taken a great interest in the subject of medical education, may entitle me to a hearing. I mean the nature of medical education itself, and the co-operation of the university in its promotion.

What is the object of medical education? It is to enable the practitioner, on the one hand, to prevent disease by his knowledge of hygiene; on the other hand, to divine its nature, and to alleviate or cure it, by his knowledge of pathology, therapeutics, and practical medicine. That is his business in life, and if he has not a thorough and practical knowledge of the conditions of health, of the causes which tend to the establishment of disease, of the meaning of symptoms, and of the uses of medicines and operative appliances, he is incompetent, even if he were the best anatomist, or physiologist, or chemist, that ever took a gold medal or won a prize certificate. This is one great truth respecting medical education. Another is, that all practice in medicine is based upon theory of some sort or other; and therefore, that it is desirable to have such theory in the closest possible accordance with fact. The veriest empiric who gives a drug in one case because he has seen it do good in another of apparently the same sort, acts upon the theory that similarity of superficial symptoms means similarity of lesions; which, by the way, is perhaps as wild an hypothesis as could be invented. To understand the nature of disease we must understand health, and the understanding of the healthy body means the having a knowledge of its structure and of the way in which its manifold actions are performed, which is what is technically termed human anatomy and human physiology. The physiologist again must needs possess an acquaintance with physics and chemistry, inasmuch as physiology is, to a great extent, applied physics and chemistry. For ordinary purposes a limited amount of such knowledge is all that is needful; but for the pursuit of the higher branches of physiology no knowledge of these branches of science can be too extensive, or too profound. Again, what we call therapeutics, which has to do with the action of drugs and medicines on the living organism, is, strictly speaking, a branch of experimental physiology, and is daily receiving a greater and greater experimental development.

The third great fact which is to be taken into consideration in dealing with medical education, is that the practical necessities of life do not, as a rule, allow aspirants to medical practice to give more than three, or it may be four years to their studies. Let us put it at four years, and then reflect that, in the course of this time, a young man fresh from school has to acquaint himself with medicine, surgery, obstetrics, therapeutics, pathology, hygiene, as well as with the anatomy and the physiology of the human body; and that his knowledge should be of such a character that it can be relied upon in any emergency, and always ready for practical application. Consider, in addition, that the medical practitioner may be called upon, at any moment, to give evidence in a court of justice in a criminal case; and that it is therefore well that he should know something of the laws of evidence, and of what we call medical jurisprudence. On a medical certificate, a man may be taken from his home and from his business and confined in a lunatic asylum; surely, therefore, it is desirable that the medical practitioner should have some rational and clear conceptions as to the nature and symptoms of mental disease. Bearing in mind all these requirements of medical education, you will admit that the burden on the young aspirant for the medical profession is somewhat of the heaviest, and that it needs some care to prevent his intellectual back from being broken.

Those who are acquainted with the existing systems of medical education will observe that, long as is the catalogue of studies which I have enumerated, I have omitted to mention several that enter into the usual medical curriculum of the present day. I have said not a word about zoology, comparative anatomy, botany, or materia medica. Assuredly this is from no light estimate of the value or importance of such studies in themselves. It may be taken for granted that I should be the last person in the world to object to the teaching of zoology, or comparative anatomy, in themselves; but I have the strongest feeling that, considering the number and the gravity of those studies through which a medical man must pass, if he is to be competent to discharge the serious duties which devolve upon him, subjects which lie so remote as these do from his practical pursuits should be rigorously excluded. The young man, who has enough to do in order to acquire such familiarity with the structure of the human body as will enable him to perform the operations of surgery, ought not, in my judgment, to be occupied with investigations into the anatomy of crabs and starfishes. Undoubtedly the doctor should know the common poisonous plants of his own country when he sees them; but that knowledge may be obtained by a few hours devoted to the examination of specimens of such plants, and the desirableness of such knowledge is no justification, to my mind, for spending three months over the study of systematic botany. Again, materia medica, so far as it is a knowledge of drugs, is the business of the druggist. In all other callings the necessity of the division of labour is fully recognised, and it is absurd to require of the medical man that he should not avail himself of the special knowledge of those whose business it is to deal in the drugs which he uses. It is all very well that the physician should know that castor oil comes from a plant, and castoreum from an animal, and how they are to be prepared; but for all the practical purposes of his profession that knowledge is not of one whit more value, has no more relevancy, than the knowledge of how the steel of his scalpel is made.

All knowledge is good. It is impossible to say that any fragment of knowledge, however insignificant or remote from one's ordinary pursuits, may not some day be turned to account. But in medical education, above all things, it is to be recollected that, in order to know a little well, one must be content to be ignorant of a great deal.

Let it not be supposed that I am proposing to narrow medical education, or, as the cry is, to lower the standard of the profession. Depend upon it there is only one way of really ennobling any calling, and that is to make those who pursue it real masters of their craft, men who can truly do that which they profess to be able to do, and which they are credited with being able to do by the public. And there is no position so ignoble as that of the so-called "liberally-educated practitioner," who, as Talleyrand said of his physician, "Knows everything, even a little physic;" who may be able to read Galen in the original; who knows all the plants, from the cedar of Lebanon to the hyssop upon the wall; but who finds himself, with the issues of life and death in his hands, ignorant, blundering, and bewildered, because of his ignorance of the essential and fundamental truths upon which practice must be based. Moreover, I venture to say, that any man who has seriously studied all the essential branches of medical knowledge; who has the needful acquaintance with the elements of physical science; who has been brought by medical jurisprudence into contact with law; whose study of insanity has taken him into the fields of psychology; has ipso facto received a liberal education.

Having lightened the medical curriculum by culling out of it everything which is unessential, we may next consider whether something may not be done to aid the medical student toward the acquirement of real knowledge by modifying the system of examination. In England, within my recollection, it was the practice to require of the medical student attendance on lectures upon the most diverse topics during three years; so that it often happened that he would have to listen, in the course of a day, to four or five lectures upon totally different subjects, in addition to the hours given to dissection and to hospital practice: and he was required to keep all the knowledge he could pick up, in this distracting fashion, at examination point, until, at the end of three years, he was set down to a table and questioned pell-mell upon all the different matters with which he had been striving to make acquaintance. A worse system and one more calculated to obstruct the acquisition of sound knowledge and to give full play to the "crammer" and the "grinder" could hardly have been devised by human ingenuity. Of late years great reforms have taken place. Examinations have been divided so as to diminish the number of subjects among which the attention has to be distributed. Practical examination has been largely introduced; but there still remains, even under the present system, too much of the old evil inseparable from the contemporaneous pursuit of a multiplicity of diverse studies.

Proposals have recently been made to get rid of general examinations altogether, to permit the student to be examined in each subject at the end of his attendance on the class; and then, in case of the result being satisfactory, to allow him to have done with it; and I may say that this method has been pursued for many years in the Royal School of Mines in London, and has been found to work very well. It allows the student to concentrate his mind upon what he is about for the time being, and then to dismiss it. Those who are occupied in intellectual work, will, I think, agree with me that it is important, not so much to know a thing, as to have known it, and known it thoroughly. If you have once known a thing in this way it is easy to renew your knowledge when you have forgotten it; and when you begin to take the subject up again, it slides back upon the familiar grooves with great facility.

Lastly comes the question as to how the university may co-operate in advancing medical education. A medical school is strictly a technical school—a school in which a practical profession is taught—while a university ought to be a place in which knowledge is obtained without direct reference to professional purposes. It is clear, therefore, that a university and its antecedent, the school, may best co-operate with the medical school by making due provision for the study of those branches of knowledge which lie at the foundation of medicine.

At present, young men come to the medical schools without a conception of even the elements of physical science; they learn, for the first time, that there are such sciences as physics, chemistry, and physiology, and are introduced to anatomy as a new thing. It may be safely said that, with a large proportion of medical students, much of the first session is wasted in learning how to learn—in familiarising themselves with utterly strange conceptions, and in awakening their dormant and wholly untrained powers of observation and of manipulation. It is difficult to overestimate the magnitude of the obstacles which are thrown in the way of scientific training by the existing system of school education. Not only are men trained in mere book-work, ignorant of what observation means, but the habit of learning from books alone begets a disgust of observation. The book-learned student will rather trust to what he sees in a book than to the witness of his own eyes.

There is not the least reason why this should be so, and, in fact, when elementary education becomes that which I have assumed it ought to be, this state of things will no longer exist. There is not the slightest difficulty in giving sound elementary instruction in physics, in chemistry, and in the elements of human physiology, in ordinary schools. In other words, there is no reason why the student should not come to the medical school, provided with as much knowledge of these several sciences as he ordinarily picks up, in the course of his first year of attendance, at the medical school.

I am not saying this without full practical justification for the statement. For the last eighteen years we have had in England a system of elementary science teaching carried out under the auspices of the Science and Art Department, by which elementary scientific instruction is made readily accessible to the scholars of all the elementary schools in the country. Commencing with small beginnings, carefully developed and improved, that system now brings up for examination as many as seven thousand scholars in the subject of human physiology alone. I can say that, out of that number, a large proportion have acquired a fair amount of substantial knowledge; and that no inconsiderable percentage show as good an acquaintance with human physiology as used to be exhibited by the average candidates for medical degrees in the University of London, when I was first an examiner there twenty years ago; and quite as much knowledge as is possessed by the ordinary student of medicine at the present day. I am justified, therefore, in looking forward to the time when the student who proposes to devote himself to medicine will come, not absolutely raw and inexperienced as he is at present, but in a certain state of preparation for further study; and I look to the university to help him still further forward in that stage of preparation, through the organisation of its biological department. Here the student will find means of acquainting himself with the phenomena of life in their broadest acceptation. He will study not botany and zoology, which, as I have said, would take him too far away from his ultimate goal; but, by duly arranged instruction, combined with work in the laboratory upon the leading types of animal and vegetable life, he will lay a broad, and at the same time solid, foundation of biological knowledge; he will come to his medical studies with a comprehension of the great truths of morphology and of physiology, with his hands trained to dissect and his eyes taught to see. I have no hesitation in saying that such preparation is worth a full year added on to the medical curriculum. In other words, it will set free that much time for attention to those studies which bear directly upon the student's most grave and serious duties as a medical practitioner.

Up to this point I have considered only the teaching aspect of your great foundation, that function of the university in virtue of which it plays the part of a reservoir of ascertained truth, so far as our symbols can ever interpret nature. All can learn; all can drink of this lake. It is given to few to add to the store of knowledge, to strike new springs of thought, or to shape new forms of beauty. But so sure as it is that men live not by bread, but by ideas, so sure is it that the future of the world lies in the hands of those who are able to carry the interpretation of nature a step further than their predecessors; so certain is it that the highest function of a university is to seek out those men, cherish them, and give their ability to serve their kind full play.

I rejoice to observe that the encouragement of research occupies so prominent a place in your official documents, and in the wise and liberal inaugural address of your president. This subject of the encouragement, or, as it is sometimes called, the endowment of research, has of late years greatly exercised the minds of men in England. It was one of the main topics of discussion by the members of the Royal Commission of whom I was one, and who not long since issued their report, after five years' labour. Many seem to think that this question is mainly one of money; that you can go into the market and buy research, and that supply will follow demand, as in the ordinary course of commerce. This view does not commend itself to my mind. I know of no more difficult practical problem than the discovery of a method of encouraging and supporting the original investigator without opening the door to nepotism and jobbery. My own conviction is admirably summed up in the passage of your president's address, "that the best investigators are usually those who have also the responsibilities of instruction, gaining thus the incitement of colleagues, the encouragement of pupils, and the observation of the public."

At the commencement of this address I ventured to assume that I might, if I thought fit, criticise the arrangements which have been made by the board of trustees, but I confess that I have little to do but to applaud them. Most wise and sagacious seems to me the determination not to build for the present. It has been my fate to see great educational funds fossilise into mere bricks and mortar, in the petrifying springs of architecture, with nothing left to work the institution they were intended to support. A great warrior is said to have made a desert and called it peace. Administrators of educational funds have sometimes made a palace and called it a university. If I may venture to give advice in a matter which lies out of my proper competency, I would say that whenever you do build, get an honest bricklayer, and make him build you just such rooms as you really want, leaving ample space for expansion. And a century hence, when the Baltimore and Ohio shares are at one thousand premium, and you have endowed all the professors you need, and built all the laboratories that are wanted, and have the best museum and the finest library that can be imagined; then, if you have a few hundred thousand dollars you don't know what to do with, send for an architect and tell him to put up a facade. If American is similar to English experience, any other course will probably lead you into having some stately structure, good for your architect's fame, but not in the least what you want.

It appears to me that what I have ventured to lay down as the principles which should govern the relations of a university to education in general, are entirely in accordance with the measures you have adopted. You have set no restrictions upon access to the instruction you propose to give; you have provided that such instruction, either as given by the university or by associated institutions, should cover the field of human intellectual activity. You have recognised the importance of encouraging research. You propose to provide means by which young men, who may be full of zeal for a literary or for a scientific career, but who also may have mistaken aspiration for inspiration, may bring their capacities to a test, and give their powers a fair trial. If such a one fail, his endowment terminates, and there is no harm done. If he succeed, you may give power of flight to the genius of a Davy or a Faraday, a Carlyle or a Locke, whose influence on the future of his fellow-men shall be absolutely incalculable.

You have enunciated the principle that "the glory of the university should rest upon the character of the teachers and scholars, and not upon their numbers or buildings constructed for their use." And I look upon it as an essential and most important feature of your plan that the income of the professors and teachers shall be independent of the number of students whom they can attract. In this way you provide against the danger, patent elsewhere, of finding attempts at improvement obstructed by vested interests; and, in the department of medical education especially, you are free of the temptation to set loose upon the world men utterly incompetent to perform the serious and responsible duties of their profession.

It is a delicate matter for a stranger to the practical working of your institutions, like myself, to pretend to give an opinion as to the organisation of your governing power. I can conceive nothing better than that it should remain as it is, if you can secure a succession of wise, liberal, honest, and conscientious men to fill the vacancies that occur among you. I do not greatly believe in the efficacy of any kind of machinery for securing such a result; but I would venture to suggest that the exclusive adoption of the method of co-optation for filling the vacancies which must occur in your body, appears to me to be somewhat like a tempting of Providence. Doubtless there are grave practical objections to the appointment of persons outside of your body and not directly interested in the welfare of the university; but might it not be well if there were an understanding that your academic staff should be officially represented on the board, perhaps even the heads of one or two independent learned bodies, so that academic opinion and the views of the outside world might have a certain influence in that most important matter, the appointment of your professors? I throw out these suggestions, as I have said, in ignorance of the practical difficulties that may lie in the way of carrying them into effect, on the general ground that personal and local influences are very subtle, and often unconscious, while the future greatness and efficiency of the noble institution which now commences its work must largely depend upon its freedom from them.

* * * * *

I constantly hear Americans speak of the charm which our old mother country has for them, of the delight with which they wander through the streets of ancient towns, or climb the battlements of mediaeval strongholds, the names of which are indissolubly associated with the great epochs of that noble literature which is our common inheritance; or with the blood-stained steps of that secular progress, by which the descendants of the savage Britons and of the wild pirates of the North Sea have become converted into warriors of order and champions of peaceful freedom, exhausting what still remains of the old Berserk spirit in subduing nature, and turning the wilderness into a garden. But anticipation has no less charm than retrospect, and to an Englishman landing upon your shores for the first time, travelling for hundreds of miles through strings of great and well-ordered cities, seeing your enormous actual, and almost infinite potential, wealth in all commodities, and in the energy and ability which turn wealth to account, there is something sublime in the vista of the future. Do not suppose that I am pandering to what is commonly understood by national pride. I cannot say that I am in the slightest degree impressed by your bigness, or your material resources, as such. Size is not grandeur, and territory does not make a nation. The great issue, about which hangs a true sublimity, and the terror of overhanging fate, is what are you going to do with all these things? What is to be the end to which these are to be the means? You are making a novel experiment in politics on the greatest scale which the world has yet seen. Forty millions at your first centenary, it is reasonably to be expected that, at the second, these states will be occupied by two hundred millions of English-speaking people, spread over an area as large as that of Europe, and with climates and interests as diverse as those of Spain and Scandinavia, England and Russia. You and your descendants have to ascertain whether this great mass will hold together under the forms of a republic, and the despotic reality of universal suffrage; whether state rights will hold out against centralisation, without separation; whether centralisation will get the better, without actual or disguised monarchy; whether shifting corruption is better than a permanent bureaucracy; and as population thickens in your great cities, and the pressure of want is felt, the gaunt spectre of pauperism will stalk among you, and communism and socialism will claim to be heard. Truly America has a great future before her; great in toil, in care, and in responsibility; great in true glory if she be guided in wisdom and righteousness; great in shame if she fail. I cannot understand why other nations should envy you, or be blind to the fact that it is for the highest interest of mankind that you should succeed; but the one condition of success, your sole safeguard, is the moral worth and intellectual clearness of the individual citizen. Education cannot give these, but it may cherish them and bring them to the front in whatever station of society they are to be found; and the universities ought to be, and may be, the fortresses of the higher life of the nation.

May the university which commences its practical activity to-morrow abundantly fulfil its high purpose; may its renown as a seat of true learning, a centre of free inquiry, a focus of intellectual light, increase year by year, until men wander hither from all parts of the earth, as of old they sought Bologna, or Paris, or Oxford.

And it is pleasant to me to fancy that, among the English students who are drawn to you at that time, there may linger a dim tradition that a countryman of theirs was permitted to address you as he has done to-day, and to feel as if your hopes were his hopes and your success his joy.

[1] Delivered at the formal opening of the Johns Hopkins University at Baltimore, U.S., September 12. The total amount bequeathed by Johns Hopkins is more than 7,000,000 dollars. The sum of 3,500,000 dollars is appropriated to a university, a like sum to a hospital, and the rest to local institutions of education and charity.



LONDON.

LECTURE ON THE STUDY OF BIOLOGY.

It is my duty to-night to speak about the study of Biology, and while it may be that there are many of my audience who are quite familiar with that study, yet as a lecturer of some standing, it would, I know by experience, be very bad policy on my part to suppose such to be extensively the case. On the contrary, I must imagine that there are many of you who would like to know what Biology is; that there are others who have that amount of information, but would nevertheless gladly hear why it should be worth their while to study Biology; and yet others, again, to whom these two points are clear, but who desire to learn how they had best study it, and, finally, when they had best study it.

I shall, therefore, address myself to the endeavour to give you some answer to these four questions—what Biology is; why it should be studied; how it should be studied; and when it should be studied.

In the first place, in respect to what Biology is, there are, I believe, some persons who imagine that the term "Biology" is simply a new-fangled denomination, a neologism in short, for what used to be known under the title of "Natural History;" but I shall try to show you, on the contrary, that the word is the expression of the growth of science during the last 200 years, and came into existence half a century ago.

At the revival of learning, knowledge was divided into two kinds—the knowledge of nature and the knowledge of man; for it was the current idea then (and a great deal of that ancient conception still remains) that there was a sort of essential antithesis, not to say antagonism, between nature and man; and that the two had not very much to do with one another, except that the one was oftentimes exceedingly troublesome to the other. Though it is one of the salient merits of our great philosophers of the seventeenth century, that they recognised but one scientific method, applicable alike to man and to nature, we find this notion of the existence of a broad distinction between nature and man in the writings both of Bacon and of Hobbes of Malmesbury; and I have brought with me that famous work which is now so little known, greatly as it deserves to be studied, "The Leviathan," in order that I may put to you in the wonderfully terse and clear language of Thomas Hobbes, what was his view of the matter. He says:—

"The register of knowledge of fact is called history. Whereof there be two sorts, one called natural history; which is the history of such facts or effects of nature as have no dependence on man's will; such as are the histories of metals, plants, animals, regions, and the like. The other is civil history; which is the history of the voluntary actions of men in commonwealths."

So that all history of fact was divided into these two great groups of natural and of civil history. The Royal Society was in course of foundation about the time that Hobbes was writing this book, which was published in 1651; and that Society was termed a "Society for the Improvement of Natural Knowledge," which was then nearly the same thing as a "Society for the Improvement of Natural History." As time went on, and the various branches of human knowledge became more distinctly developed and separated from one another, it was found that some were much more susceptible of precise mathematical treatment than others. The publication of the "Principia" of Newton, which probably gave a greater stimulus to physical science than any work ever published before, or which is likely to be published hereafter, showed that precise mathematical methods were applicable to those branches of science such as astronomy, and what we now call physics, which occupy a very large portion of the domain of what the older writers understood by natural history. And inasmuch as the partly deductive and partly experimental methods of treatment to which Newton and others subjected these branches of human knowledge, showed that the phenomena of nature which belonged to them were susceptible of explanation, and thereby came within the reach of what was called "philosophy" in those days; so much of this kind of knowledge as was not included under astronomy came to be spoken of as "natural philosophy"—a term which Bacon had employed in a much wider sense. Time went on, and yet other branches of science developed themselves. Chemistry took a definite shape; and since all these sciences, such as astronomy, natural philosophy, and chemistry, were susceptible either of mathematical treatment or of experimental treatment, or of both, a broad distinction was drawn between the experimental branches of what had previously been called natural history and the observational branches—those in which experiment was (or appeared to be) of doubtful use, and where, at that time, mathematical methods were inapplicable. Under these circumstances the old name of "Natural History" stuck by the residuum, by those phenomena which were not, at that time, susceptible of mathematical or experimental treatment; that is to say, those phenomena of nature which come now under the general heads of physical geography, geology, mineralogy, the history of plants, and the history of animals. It was in this sense that the term was understood by the great writers of the middle of the last century—Buffon and Linnaeus—by Buffon in his great work, the "Histoire Naturelle Generale," and by Linnaeus in his splendid achievement, the "Systema Naturae." The subjects they deal with are spoken of as "Natural History," and they called themselves and were called "Naturalists." But you will observe that this was not the original meaning of these terms; but that they had, by this time, acquired a signification widely different from that which they possessed primitively.

The sense in which "Natural History" was used at the time I am now speaking of has, to a certain extent, endured to the present day. There are now in existence in some of our northern universities, chairs of "Civil and Natural History," in which "Natural History" is used to indicate exactly what Hobbes and Bacon meant by that term. The unhappy incumbent of the chair of Natural History is, or was, supposed to cover the whole ground of geology, mineralogy, and zoology, perhaps even botany, in his lectures.

But as science made the marvellous progress which it did make at the latter end of the last and the beginning of the present century, thinking men began to discern that under this title of "Natural History" there were included very heterogeneous constituents—that, for example, geology and mineralogy were, in many respects, widely different from botany and zoology; that a man might obtain an extensive knowledge of the structure and functions of plants and animals, without having need to enter upon the study of geology or mineralogy, and vice versa; and, further as knowledge advanced, it became clear that there was a great analogy, a very close alliance, between those two sciences of botany and zoology which deal with living beings, while they are much more widely separated from all other studies. It is due to Buffon to remark that he clearly recognised this great fact. He says: "Ces deux genres d'etres organises [les animaux et les vegetaux] ont beaucoup plus de proprietes communes que de differences reelles." Therefore, it is not wonderful that, at the beginning of the present century, in two different countries, and so far as I know, without any intercommunication, two famous men clearly conceived the notion of uniting the sciences which deal with living matter into one whole, and of dealing with them as one discipline. In fact, I may say there were three men to whom this idea occurred contemporaneously, although there were but two who carried it into effect, and only one who worked it out completely. The persons to whom I refer were the eminent physiologist Bichat, and the great naturalist Lamarck, in France; and a distinguished German, Treviranus. Bichat[1] assumed the existence of a special group of "physiological" sciences. Lamarck, in a work published in 1801,[2] for the first time made use of the name "Biologie" from the two Greek words which signify a discourse upon life and living things. About the same time it occurred to Treviranus, that all those sciences which deal with living matter are essentially and fundamentally one, and ought to be treated as a whole; and, in the year 1802, he published the first volume of what he also called "Biologie." Treviranus's great merit lies in this, that he worked out his idea, and wrote the very remarkable book to which I refer. It consists of six volumes, and occupied its author for twenty years—from 1802 to 1822.

That is the origin of the term "Biology;" and that is how it has come about that all clear thinkers and lovers of consistent nomenclature have substituted for the old confusing name of "Natural History," which has conveyed so many meanings, the term "Biology" which denotes the whole of the sciences which deal with living things, whether they be animals or whether they be plants. Some little time ago—in the course of this year, I think—I was favoured by a learned classic, Dr. Field of Norwich, with a disquisition, in which he endeavoured to prove that, from a philological point of view, neither Treviranus nor Lamarck had any right to coin this new word "Biology" for their purpose; that, in fact, the Greek word "Bios" had relation only to human life and human affairs, and that a different word was employed by the Greeks when they wished to speak of the life of animals and plants. So Dr. Field tells us we are all wrong in using the term biology, and that we ought to employ another; only he is not quite sure about the propriety of that which he proposes as a substitute. It is a somewhat hard one—"zootocology." I am sorry we are wrong, because we are likely to continue so. In these matters we must have some sort of "Statute of Limitations." When a name has been employed for half-a-century, persons of authority[3] have been using it, and its sense has become well understood, I am afraid that people will go on using it, whatever the weight of philological objection.

Now that we have arrived at the origin of this word "Biology," the next point to consider is: What ground does it cover? I have said that, in its strict technical sense, it denotes all the phenomena which are exhibited by living things, as distinguished from those which are not living; but while that is all very well, so long as we confine ourselves to the lower animals and to plants, it lands us in considerable difficulties when we reach the higher forms of living things. For whatever view we may entertain about the nature of man, one thing is perfectly certain, that he is a living creature. Hence, if our definition is to be interpreted strictly, we must include man and all his ways and works under the head of Biology; in which case, we should find that psychology, politics, and political economy would be absorbed into the province of Biology. In fact, civil history would be merged in natural history. In strict logic it may be hard to object to this course, because no one can doubt that the rudiments and outlines of our own mental phenomena are traceable among the lower animals. They have their economy and their polity, and if, as is always admitted, the polity of bees and the commonwealth of wolves fall within the purview of the biologist proper, it becomes hard to say why we should not include therein human affairs, which in so many cases resemble those of the bees in zealous getting, and are not without a certain parity in the proceedings of the wolves. The real fact is that we biologists are a self-sacrificing people; and inasmuch as, on a moderate estimate, there are about a quarter of a million different species of animals and plants to know about already, we feel that we have more than sufficient territory. There has been a sort of practical convention by which we give up to a different branch of science what Bacon and Hobbes would have called "Civil History." That branch of science has constituted itself under the head of Sociology. I may use phraseology which, at present, will be well understood and say that we have allowed that province of Biology to become autonomous; but I should like you to recollect that that is a sacrifice, and that you should not be surprised if it occasionally happens that you see a biologist apparently trespassing in the region of philosophy or politics; or meddling with human education; because, after all, that is a part of his kingdom which he has only voluntarily forsaken.

Having now defined the meaning of the word Biology, and having indicated the general scope of Biological Science, I turn to my second question, which is—Why should we study Biology? Possibly the time may come when that will seem a very odd question. That we, living creatures, should not feel a certain amount of interest in what it is that constitutes our life will eventually, under altered ideas of the fittest objects of human inquiry, appear to be a singular phenomenon; but, at present, judging by the practice of teachers and educators, Biology would seem to be a topic that does not concern us at all. I propose to put before you a few considerations with which I dare say many will be familiar already, but which will suffice to show—not fully, because to demonstrate this point fully would take a great many lectures—that there are some very good and substantial reasons why it may be advisable that we should know something about this branch of human learning.

I myself entirely agree with another sentiment of the philosopher of Malmesbury, "that the scope of all speculation is the performance of some action or thing to be done," and I have not any very great respect for, or interest in, mere knowing as such. I judge of the value of human pursuits by their bearing upon human interests; in other words, by their utility; but I should like that we should quite clearly understand what it is that we mean by this word "utility." In an Englishman's mouth it generally means that by which we get pudding or praise, or both. I have no doubt that is one meaning of the word utility, but it by no means includes all I mean by utility. I think that knowledge of every kind is useful in proportion as it tends to give people right ideas, which are essential to the foundation of right practice, and to remove wrong ideas, which are the no less essential foundations and fertile mothers of every description of error in practice. And inasmuch as, whatever practical people may say, this world is, after all, absolutely governed by ideas, and very often by the wildest and most hypothetical ideas, it is a matter of the very greatest importance that our theories of things, and even of things that seem a long way apart from our daily lives, should be as far as possible true, and as far as possible removed from error. It is not only in the coarser practical sense of the word "utility," but in this higher and broader sense, that I measure the value of the study of biology by its utility; and I shall try to point out to you that you will feel the need of some knowledge of biology at a great many turns of this present nineteenth century life of ours. For example, most of us attach great importance to the conception which we entertain of the position of man in this universe and his relation to the rest of nature. We have almost all been told, and most of us hold by the tradition, that man occupies an isolated and peculiar position in nature; that though he is in the world he is not of the world; that his relations to things about him are of a remote character; that his origin is recent, his duration likely to be short, and that he is the great central figure round which other things in this world revolve. But this is not what the biologist tells us.

At the present moment you will be kind enough to separate me from them, because it is in no way essential to my present argument that I should advocate their views. Don't suppose that I am saying this for the purpose of escaping the responsibility of their beliefs; indeed, at other times and in other places, I do not think that point has been left doubtful; but I want clearly to point out to you that for my present argument they may all be wrong; and, nevertheless, my argument will hold good. The biologists tell us that all this is an entire mistake. They turn to the physical organisation of man. They examine his whole structure, his bony frame and all that clothes it. They resolve him into the finest particles into which the microscope will enable them to break him up. They consider the performance of his various functions and activities, and they look at the manner in which he occurs on the surface of the world. Then they turn to other animals, and taking the first handy domestic animal—say a dog—they profess to be able to demonstrate that the analysis of the dog leads them, in gross, to precisely the same results as the analysis of the man; that they find almost identically the same bones, having the same relations; that they can name the muscles of the dog by the names of the muscles of the man, and the nerves of the dog by those of the nerves of the man, and that, such structures and organs of sense as we find in the man such also we find in the dog; they analyse the brain and spinal cord, and they find that the nomenclature which fits the one answers for the other. They carry their microscopic inquiries in the case of the dog as far as they can, and they find that his body is resolvable into the same elements as those of the man. Moreover, they trace back the dog's and the man's development, and they find that, at a certain stage of their existence, the two creatures are not distinguishable the one from the other; they find that the dog and his kind have a certain distribution over the surface of the world, comparable in its way to the distribution of the human species. What is true of the dog they tell us is true of all the higher animals; and they assert that they can lay down a common plan for the whole of these creatures, and regard the man and the dog, the horse and the ox as minor modifications of one great fundamental unity. Moreover, the investigations of the last three-quarters of a century have proved, they tell us, that similar inquiries, carried out through all the different kinds of animals which are met with in nature, will lead us, not in one straight series, but by many roads, step by step, gradation by gradation, from man, at the summit, to specks of animated jelly at the bottom of the series. So that the idea of Leibnitz, and of Bonnet, that animals form a great scale of being, in which there are a series of gradations from the most complicated form to the lowest and simplest; that idea, though not exactly in the form in which it was propounded by those philosophers, turns out to be substantially correct. More than this, when biologists pursue their investigations into the vegetable world, they find that they can, in the same way, follow out the structure of the plant, from the most gigantic and complicated trees down through a similar series of gradations, until they arrive at specks of animated jelly, which they are puzzled to distinguish from those specks which they reached by the animal road.

Thus, biologists have arrived at the conclusion that a fundamental uniformity of structure pervades the animal and vegetable worlds, and that plants and animals differ from one another simply as diverse modifications of the same great general plan.

Again, they tell us the same story in regard to the study of function. They admit the large and important interval which, at the present time, separates the manifestations of the mental faculties observable in the higher forms of mankind, and even in the lower forms, such as we know them, from those exhibited by other animals; but, at the same time, they tell us that the foundations, or rudiments, of almost all the faculties of man are to be met with in the lower animals; that there is a unity of mental faculty as well as of bodily structure, and that, here also, the difference is a difference of degree and not of kind. I said "almost all," for a reason. Among the many distinctions which have been drawn between the lower creatures and ourselves, there is one which is hardly ever insisted on,[4] but which may be very fitly spoken of in a place so largely devoted to Art as that in which we are assembled. It is this, that while, among various kinds of animals, it is possible to discover traces of all the other faculties of man, especially the faculty of mimicry, yet that particular form of mimicry which shows itself in the imitation of form, either by modelling or by drawing, is not to be met with. As far as I know, there is no sculpture or modelling, and decidedly no painting or drawing, of animal origin, I mention the fact, in order that such comfort may be derived therefrom as artists may feel inclined to take.

If what the biologists tell us is true, it will be needful to get rid of our erroneous conceptions of man, and of his place in nature, and to substitute right ones for them. But it is impossible to form any judgment as to whether the biologists are right or wrong, unless we are able to appreciate the nature of the arguments which they have to offer.

One would almost think this to be a self-evident proposition. I wonder what a scholar would say to the man who should undertake to criticise a difficult passage in a Greek play, but who obviously had not acquainted himself with the rudiments of Greek grammar. And yet, before giving positive opinions about these high questions of Biology, people not only do not seem to think it necessary to be acquainted with the grammar of the subject, but they have not even mastered the alphabet. You find criticism and denunciation showered about by persons, who, not only have not attempted to go through the discipline necessary to enable them to be judges, but who have not even reached that stage of emergence from ignorance in which the knowledge that such a discipline is necessary dawns upon the mind. I have had to watch with some attention—in fact I have been favoured with a good deal of it myself—the sort of criticism with which biologists and biological teachings are visited. I am told every now and then that there is a "brilliant article"[5] in so-and-so, in which we are all demolished. I used to read these things once, but I am getting old now, and I have ceased to attend very much to this cry of "wolf." When one does read any of these productions, what one finds generally, on the face of it, is that the brilliant critic is devoid of even the elements of biological knowledge, and that his brilliancy is like the light given out by the crackling of thorns under a pot of which Solomon speaks. So far as I recollect, Solomon makes use of the image for purposes of comparison; but I will not proceed further into that matter.

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